Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
A few examples of _Phyllotheca deliquescens_ have been described by
Schmalhausen in which a branch bears clusters of sporangiophores,
alternating with verticils of sterile bracts. The sporangiophores
appear to have the form of stalked peltate appendages bearing
sporangia, very similar to the sporangiophores of _Equisetum_.
Solms-Laubach[561] has examined the best of Schmalhausen’s specimens,
and a carefully drawn figure of one of the fertile branches is given in
his _Fossil Botany_.
The significance of this manner of occurrence of sporangiophores
and whorls of sterile bracts on the fertile branch will be better
understood after a description of the strobilus of _Calamites_. In
_Phyllotheca_ the sporangiophores appear to have been given off in
whorls, which were separated from one another by whorls of sterile
bracts, whereas in _Equisetum_ there are no sterile appendages
associated with the sporangiophores of the strobilus, with the
exception of the annulus at the base of the cone. Heer[562] first drew
attention to the fact that in _Phyllotheca_ we have a form of strobilus
or fertile shoot to a certain extent intermediate in character between
_Equisetum_ and _Calamites_.
In abnormal fertile shoots of _Equisetum_, sporophylls occasionally
occur above and below a sterile leaf-sheath. Potonié[563] has figured
such an example in which an apical strobilus is succeeded at a lower
level by a sterile leaf-sheath, and this again by a second cluster
of sporophylls. As Potonié points out, this alternation of fertile
and sterile members affords an interesting resemblance between
_Phyllotheca_ and _Equisetum_. It suggests a partial reversion towards
the Calamitean type of strobilus.
2. _Phyllotheca Brongniarti_ Zigno. Fig. 68, A.
This species of _Phyllotheca_ from the Lower Oolite rocks of Italy
is known only in the form of sterile branches. The leaves are fused
basally into an open cup-like sheath which is dissected into several
spreading and incurved linear segments. The internodes are striated
longitudinally; they are about 2 mm. in diameter and 10 mm. in length.
The specimen represented in fig. 68, A, was originally described by
the Italian palaeobotanist Zigno[564]; it serves to illustrate the
points of difference between this genus and the ordinary _Equisetum_.
The open and spreading sheaths clasping the nodes and the erect
solitary branches give the plant a distinctive appearance.
[Illustration: FIG. 68.
A. _Phyllotheca Brongniarti_, Zigno. Nat. size. (After Zigno.)
B. _Calamocladus frondosus_, Grand’Eury. (After Grand’Eury.)
Slightly enlarged.
C. _Phyllotheca indica_, Bunb. Part of a leaf-sheath. From a specimen
in the Museum of the Geological Society. Slightly enlarged.]
3. _Phyllotheca indica_ Bunb. and _P. australis_ Brongn. Fig. 68, C.
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