Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
In the small portion of a stem represented in fig. 74 _B_, the cortical
tissues have been partially preserved; at the inner edge, next the
hollow pith, there are two xylem groups, each with a carinal canal, and
between them is part of a broad “principal” medullary ray[614]. The
cambium has not been preserved, but beyond this region we have some
of the large cells, _c_, of the inner cortex; these are followed by a
few remnants of a smaller-celled tissue, and external to this part of
the cortex there is a series of triangular groups, _h_, consisting of
small thick-walled cells alternating with spaces which were originally
occupied by more delicate parenchyma. The darker groups constitute
hypodermal strands of mechanical tissue or stereome which lent support
to the stem. The surface of a stem possessing such supporting strands
would probably assume a longitudinally wrinkled or grooved appearance
on drying; the intervening parenchyma, contracting and yielding more
readily, would tend to produce shallow grooves alternating with the
ridges above the stereome strands.
The complete section of the stem of which a small portion is shown
in fig. 74 _B_, is figured by Williamson[615] in his 12th memoir on
Coal-Measure plants. The section was obtained from Ashton-under-Lyne
in Lancashire; it illustrates very clearly a method of preservation
which is occasionally met with among petrified plants. The walls of the
various tissue elements are black in colour and somewhat ragged, and
the general appearance of the section is similar to that of a section
of a charred piece of stem. It is possible that the Calamite twig was
reduced to charcoal before petrifaction by a lightning flash or some
other cause.
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