Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The most characteristic feature of _Calamodendron_ is the occurrence of
several rows of radially disposed thick-walled prosenchymatous elements
(fig. 84, _b_) on either flank of each wedge-shaped group of xylem.
Each principal ray is thus nearly filled up by bands of fibrous cells
on the sides of adjacent xylem groups, but the centre of each principal
ray is occupied by a narrow band of parenchyma (fig. 84, _c_). The
relative breadth of the xylem and prosenchymatous bands has been made
use of by Renault as a specific character in _Calamodendron_ stems.
Fig. 84 is copied from a drawing recently published by this French
author of a new species of _Calamodendron_, _C. intermedium_[635]. In
this case the bands of fibrous cells, _b_, are slightly broader, as
seen in a transverse section of the stem, than the bands of xylem
tracheids, _a_. The narrow band, _c_, consists of four rows of the
parenchymatous tissue of a medullary ray. At the inner end of each
group of tracheids there is a large carinal canal.
The question of the recognition of the pith-casts of stems possessing
the structure of any of the three subgenera of _Calamites_ is referred
to in a later section of this chapter.
_b._ _Leaves_
_Leaves of Calamites and Calamitean foliage-shoots, including an
account of_ (α) Calamocladus (Asterophyllites) _and_ (β)
Annularia.
Our knowledge of the structure and manner of occurrence of Calamite
leaves is very incomplete. There are numerous foliage-shoots among the
fossils of the Coal-Measures which are no doubt Calamitean, but as
they are nearly always found apart from the main branches and stems,
it is generally impossible to do more than speak of them as probably
the leaf-bearing branches of a Calamite. The familiar fossils known
as _Asterophyllites_, and in recent years often referred to the genus
_Calamocladus_, are no doubt Calamitean shoots; but they are usually
found as isolated fragments, and it is seldom that we are able to refer
them to definite forms of _Calamites_. Another common Coal-Measure
genus, _Annularia_, is also Calamitean, and at least some of the
species are no doubt leafy shoots of _Calamites_. Although it is
generally accepted that the fossils referred to as _Asterophyllites_
or _Calamocladus_ are portions of _Calamites_, and not distinct
plants, it is convenient, and indeed necessary, to retain such a term
as _Calamocladus_ as a means of recording foliage-shoots, which may
possess both a botanical and a geological value.
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