Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
In the works of earlier writers we find frequent illustrations of
specimens of _Annularia_, which are compared with Asters and other
recent flowering plants. Lehmann[658] contributed a paper to the Royal
Academy of Berlin in 1756, in which he referred to certain fossil
plants as probable examples of flowers, among them being a specimen
of _Annularia_. He refers to the occurrence of fossil ferns and other
plants, and asks why we do not find flowers of the rose or tulip;
his object being “not to acquire vain glory, but to give occasion for
others to look into the matter more clearly.”
The general habit of the fossils which are now included under
_Annularia_ agrees closely with that of _Calamocladus_. There is the
same spreading form and a similar foliage in the two genera, but in
_Annularia_ the members of a whorl are always fused into a basal
sheath, and the segments are not of equal length. We may thus summarise
the characteristic features of the genus:—
Opposite branches are given off in one plane from the nodes of a
main axis; the leaves are in the form of narrow sheaths divided into
numerous and unequal linear or narrow lanceolate segments, each with
a median vein. The segments in each whorl appear to be spread out in
one plane very oblique to the axis of a branch, instead of spreading
radially in all directions; the lateral segments are usually longer
than the upper and lower members of a whorl. The vegetative branches
possess the same type of structure as _Calamites_.
A comparison of _Annularia_ and _Phyllotheca_ has already been
made in Chapter IX. (p. 282). Potonié[659] has recently given a
detailed account of Annularian leaves; he compares them with those
of _Equisetum_, and describes the occurrence on the lamina of each
leaf-segment of a broad central band or midrib, with a groove, probably
containing stomata, on either side. He shows that in well-preserved
specimens of _Annularia_, it is possible to recognise certain minute
surface-features, such as the presence of hairs and stomata, which
enable one to detect a close resemblance between the leaves of Calamite
stems and those of Annularian shoots.
It is not always easy to distinguish between _Annularia_ and
_Calamocladus_; the collar-like basal sheath in the leaves of the
former is a characteristic feature, but that cannot always be
recognised. On the other hand, the leaves of _Calamocladus_ may
sometimes be flattened out on the surface of the rock and simulate the
deeply cut sheaths of _Annularia_. It is difficult to decide how far
the manner of occurrence of Annularian leaves in one plane, which is
commonly insisted on as a generic character, is an original feature, or
how far it is the result of compression in fossilisation. Probably the
leaves of a living _Annularia_ were spread out at right angles to the
axis, as in the ‘verticils’ of such a plant as _Galium_.
Public-domain text, read in full here on John Shaqi.
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