Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
Annularian branches are occasionally found with cones given off
from the axils of some of the leaf-whorls. An interesting specimen,
which is now in the Leipzig Museum, was described by Sterzel in
1882[677], showing cones attached to a vegetative shoot of _Annularia
sphenophylloides_. The long and narrow strobili—2·5 cm. long and
about 6 mm. broad—appear very large in proportion to the size of the
vegetative branches. A fertile shoot consists of a central axis bearing
whorls of bracts alternating with sporangiophores, to each of which
are attached four sporangia. The specimen in fig. 89, _A_, does not
show the details clearly; each transverse constriction represents the
attachment of a whorl of linear bracts; the whole cone appears to
consist of a series of short broad segments. The divisions in the lower
half of each segment mark the position of the sterile bracts, while
those of the upper half represent the outlines of the upper sporangia
of each whorl of sporangiophores, the lower sporangia being hidden by
the ring of linear bracts[678]. On some portions of the specimen of
fig. 89, _A_, it is possible to recognise the outlines of cells on
the coaly surface-film; these probably belong to the sporangium wall.
This type of cone is included under the genus _Calamostachys_, a name
applied to Calamitean strobili with certain morphological characters,
as described on p. 351.
_c. Roots._
In 1871 Williamson[679] described some sections of what he considered
to be a distinct variety of a Calamite stem. The chief peculiarity
which he noticed lay in the absence of carinal canals, and in the solid
pith. Some years later the same observer[680] came to the conclusion
that the specimens were probably those of a plant generically distinct
from _Calamites_; he accordingly proposed a new name _Astromyelon_.
Subsequently Cash and Hick[681] gave an account of some examples
of apparently another form of plant, to which they gave the name
_Myriophylloides Williamsonis_; and Williamson[682] suggested the term
_Helophyton_ as a more suitable generic designation. It was, however,
demonstrated by Spencer[683] that the plant described by Cash and Hick
was identical with Williamson’s _Astromyelon_. Williamson[684] then
gave an account of several specimens of this type illustrating various
stages in the growth and development of the _Astromyelon_ ‘stems,’
which he compared with the rhizome of the recent genus _Marsilia_.
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