Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The occurrence of fossil plants in the form of isolated fragments is a
constant source of difficulty, and is well illustrated by the numerous
examples of strobili which cannot be connected with their parent
stems. We are, however, usually able to recognise Calamitean cones
if the impressions or petrified specimens are fairly well preserved,
but it is seldom possible to correlate particular types of cones with
the corresponding species of foliage-shoots or stems. Palaeobotanical
literature contains numerous illustrations and descriptions of
long and narrow strobili designated by different generic terms such
as _Volkmannia_, _Brukmannia_, _Calamostachys_, _Macrostachya_ and
others; many of these have since been recognised as the cones of
_Calamites_, while some species of _Volkmannia_ have been identified
with _Sphenophyllum_ stems. Before further considering the general
question of Calamite cones, a few examples may be described in detail
as types of fructification which are known to have been borne by
_Calamites_. The examples selected are species of the two provisional
genera _Calamostachys_ and _Palaeostachya_.
The usual form of a Calamite cone is illustrated in fig. 93, which
represents a fertile shoot bearing a few narrow linear leaves of the
_Calamocladus_ type; in the axils of some of these are borne the long
strobili.
[Illustration: FIG. 93. _Calamostachys_ sp. A fertile Calamitean shoot.
From a specimen in the Geological Survey Museum, Jermyn Street,
London. From the Upper Coal-Measures of Monmouthshire (No. 5539).]
_Calamostachys Binneyana_ (Carr.). Figs. 94 and 95.
In 1867 Carruthers[702] gave an account of the structural features
of the species of cones named by him _Volkmannia Ludwigi_ and
_V. Binneyi_, the generic term having been originally used by
Sternberg[703] for some impressions of Carboniferous strobili.
Brongniart[704] in 1849 referred to the various forms of _Volkmannia_
as cones of Asterophyllitean branches, and the latter he regarded
as the foliage-shoots of a Calamite stem. In 1868 Binney[705]
published a description, with several illustrations, of the cones
named by Carruthers _Volkmannia Binneyi_, and referred to them
as the fructification of that type of Calamite stem spoken of
in a previous section of this chapter (p. 311) as _Calamites_
(_Arthropitys_) _communis_ (Binney). This cone is now usually spoken
of as _Calamostachys Binneyana_; the specific name _Binneyana_
being suggested by Schimper[706] in 1869 as more euphonious than
that proposed by Carruthers. In recent years our knowledge of both
_C. Binneyana_ and _C. Ludwigi_ has been considerably extended. We
shall confine our attention in the following account to the former
species[707]. Some excellent figures of the latter species may be found
in Weiss’ Memoir[708] on Calamarieae.
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