Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The axis of the cone is occupied by a single stele, usually triangular
in section; the stele consists of a solid pith of elongated cells
surrounded by six vascular bundles, two at each corner. A somewhat
irregular gap marks the position of the protoxylem of each strand,
and portions of spiral or annular tracheids may occasionally be
seen in the cavity. These cavities, which may be spoken of as the
carinal canals, disappear at the nodes, where there is a mass of
short reticulately pitted tracheids, as in a Calamite stem. Vascular
bundles pass upwards in an oblique direction from the central stele
to supply the bracts, each of which is traversed by a single strand
of tracheids. The coherent portion, or disc, of each sterile whorl
consists of sclerenchymatous elements towards the upper surface, and
of parenchyma below. The pedicel of the sporangiophores consists of
fairly thick-walled cells traversed by a single vascular strand,
and the peltate distal portions are made up of parenchymatous cells
arranged in a palisade-like form at right angles to the free surface
of the sporangiophores. The vascular strand of the pedicel forks into
two halves just below the peltate head, and these branches again
bifurcate to send a branch to each sporangium. The four sporangia of
each sporangiophore are attached by a narrow band of tissue to the
shield-shaped distal expansion (fig. 95, A).
In a tangential section of a cone, such as the lower portion of fig. 94
and in fig. 95, B, the sporangiophores present the appearance of narrow
stalks (fig. 95, B, _a_) in the middle of a cluster of sporangia,
and the latter appear more or less square in outline. The wall of a
sporangium is made of a single layer of cells (fig. 95, B) which
present a characteristic appearance in surface-view (fig. 95, C), the
thin walls being crossed at right angles by small vertical plates. In
the tangential section of the coherent sterile whorls (fig. 95, B,
_b_ and _b_) the vascular strands are occasionally seen in transverse
section (fig. 95, B, _t_), as they pass outwards to the several free
bracts.
[Illustration: FIG. 95. _Calamostachys Binneyana_ (Carr.).
_A._ A sporangiophore and one sporangium. _t_, vascular bundle. × 45.
_B._ Tangential section showing portions of two sterile discs, _b_,
_b_; a sporangiophore, _a_, with its four sporangia, in two of
which are seen the spores; _t_, vascular bundle. × 35.
_C._ Surface-view of cells of a sporangium wall. × 130.
_D._ Spores and remains of mother-cells. × 130.
(After Williamson and Scott.)]
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