Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
A transverse section of the axis of the cone in the region of the
sterile and fertile appendages shows the vascular bundles arranged
in pairs. In a section through the peduncle of the cone, below the
lowest whorl of bracts, the bundles of the stele are situated at equal
distances apart. The cortical tissue of the peduncle is traversed by
a ring of large canals[721] similar to the vallecular canals of an
Equisetum stem.
Isospory is not a constant characteristic of _Palaeostachya_; some
forms have been found with macrospores and microspores[722].
_Other Calamitean cones, and examples illustrating the
connection between Cones and Vegetative Shoots._
It would be out of place in an introduction to Palaeobotany to attempt
an exhaustive account of the various cones which were probably borne
by Calamitean plants, but there are a few general points to which the
attention of the student should be directed. The examples dealt with
in the foregoing description illustrate the fact, that plants included
under the comprehensive genus _Calamites_ bore cones possessing
distinct morphological features. There are, however, other types of
strobili which have been found in organic connection with _Calamites_;
and some of these must be taken into account in dealing with
Calamarian plants. The genera _Volkmannia_, _Brukmannia_, _Huttonia_,
_Macrostachya_, in addition to _Calamostachys_ and _Palaeostachya_ and
others, have been applied by different writers to Calamitean cones. As
Solms-Laubach[723] has suggested, it is wiser to discard _Volkmannia_
and _Brukmannia_, as they have been made to do duty for cones of widely
different forms. It is better to adhere to the provisional generic
names used by Weiss, as they enable us to conveniently systematise the
various Calamarian strobili.
The following classification may be given of the better known cones,
some of which we are able to describe in considerable detail, while
others are still very imperfectly known. We have good evidence that
all these strobili were borne by vegetative shoots of the type of
_Calamites_, _Calamocladus_ or _Annularia_.
1. _Calamostachys_[724] (including _Paracalamostachys_ and
_Stachannularia_).
Cones long and narrow, consisting of a central axis bearing alternate
whorls of sterile and fertile appendages, the latter having the form of
sporangiophores attached at right angles to the axis midway between the
sterile verticils, and bearing four sporangia on the inner face of a
peltate distal expansion.
_Calamostachys Binneyana_ Schimp., _C. Ludwigi_ Carr., _C. Casheana_
Will., may be referred to as examples of this type of cone; also
some of the strobili described by different authors as species of
_Volkmannia_[725], _Brukmannia_[726], &c.
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