Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The result of the activity of this cambium band is the production of
secondary parenchyma and tracheal tissue. In fig. 179, E, drawn from
a portion of the section represented in fig. 168, B, a projecting arm
of primary xylem is seen at _x_; this is followed by 2–3 layers of
parenchymatous cells, some of which have dark contents, and beyond
this is seen a group of secondary elements, _tr_, cut across somewhat
obliquely, which are evidently products of the cambial cells on the
inner margin of the secretory zone, _sc_. The longitudinal section
(fig. 169, D) shows the cambial cells, a, next the secretory zone,
_sc_, passing internally into crushed and imperfectly preserved
elongated elements which are presumably miniature tracheae, and these
are succeeded by older and more completely lignified xylem elements,
_x_. In larger shoots the amount of secondary tissue is considerably
greater; it may consist almost entirely of short-celled parenchyma
(fig. 168, C, from _x_ to _sc_), or it may include a large proportion
of radially disposed and vertically elongated tracheae (fig. 168,
D, _x_², and fig. 170, A, _x_²), or it may consist of parenchyma
containing scattered groups of tracheae (fig. 169, A, _x_²)[326].
[Illustration: FIG. 169. _Lepidodendron fuliginosum._
A, B. (Manchester University Collection. No. Q. 645 A.)
B, C. (Manchester. No. 257.)
D. (Manchester. No. 6.)]
Fig. 169, A, is a diagrammatic sketch of the tissues—1 mm. wide—between
the primary xylem, _x_, and the inner cortex. The primary xylem is
succeeded by short parenchymatous cells followed by a zone of radially
elongated elements passing occasionally into rows of narrow scalariform
tracheae, some of which, owing to their sinuous longitudinal course
(fig. 171, C), are seen in oblique section, as at C, fig. 169, A. At
its outer edge this secondary tissue, _x_², consisting of parenchyma
and tracheae, passes into the cambial band (fig. 169, B, _a_).
[Illustration: FIG. 170. _Lepidodendron fuliginosum._ (From sections in
the Manchester Museum.)]
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