Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
Certain sections exhibiting this type of structure were described by
Binney in 1872 as _Halonia regularis_[328] on evidence supplied by Mr
Dawes, who stated that they were cut from a specimen bearing Halonia
tubercles. The section represented in fig. 172 is no doubt from an
Halonia axis. In 1890 Cash and Lomax[329] stated that they had in
their possession a stem of the _L. fuliginosum_ type with the external
features of _Lepidophloios_; this identification has been confirmed by
Kidston[330] and Weiss[331]. It is, however, equally clear that certain
species with the elongated leaf-cushions of _Lepidodendron_ must be
included among examples of shoots with the anatomical characters of _L.
fuliginosum_.
[Illustration: FIG. 173. _Lepidodendron obovatum._ (From a specimen
lent by Dr D. H. Scott.)]
Dr Scott[332] published in 1906 a short account of the structure
of a specimen from the Lower Coal-Measures of Lancashire, the
external features of which were identified by Kidston with those of
_Lepidodendron obovatum_ Sternb. Dr Scott generously allowed me to have
drawings made from his specimen; these are reproduced in fig. 173. The
form of the leaf-cushion is by no means perfect; there is a well-marked
median ridge, and the small circular scar near the upper end of some
of the cushions may represent the ligular cavity. At the base of the
leaf-cushions a cortical meristem has produced a zone of secondary
cortex; at _c_ a second meristem is seen in the outer cortex: the dark
dots in the cortex mark the positions of leaf-trace bundles. The inner
cortex, _d_, is a more compact tissue surrounding the imperfectly
preserved secretory zone. From the medullated stele a lateral branch,
_b_, is being given off; its crescentic form becoming changed to
circular as it passes nearer to the surface.
[Illustration: FIG. 174. _Lepidodendron aculeatum._
(Cambridge Botany School.)]
[Illustration: FIG. 175. _Lepidodendron aculeatum._
(Cambridge Botany School.)]
A type of _Lepidodendron_, _L. Hickii_, founded on anatomical
characters by Mr Watson[333], is believed by him to possess
leaf-cushions like those of _L. obovatum_; if this is so, it is
interesting, as he points out, to find two distinct anatomical types
associated with one species. Watson thinks it probable that the
“species” _L. obovatum_ includes at least two widely different species.
This merely emphasizes the importance of correlating structure and
external characters as far as available data permit.
[Illustration: FIG. 176. _Lepidodendron aculeatum._
(Cambridge Botany School.)]
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