Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The occurrence of long simple or branched and sometimes capitate hairs
is a common feature of Carboniferous megaspores (fig. 191, E, F, I).
It is possible that these appendages served to catch the microspores,
thus facilitating fertilisation. A peculiar form of megaspore has
been described by Mrs Scott[437], and assigned by her to _Lepidostrobus
foliaceus_, the megasporangium of which apparently contained only
four spores. As shown in fig. 191, G, a large bladder-like appendage
characterised by radiating veins is attached to the thick spore-coat;
it is suggested that this excrescence may be compared with the
“swimming” apparatus of the recent water-fern _Azolla_. The epithet
swimming which it is customary to apply to the appendages of _Azolla_
megaspores would seem to be inappropriate if Campbell[438] is correct in
stating that spores of _Azolla_ are incapable of floating.
B. _Spencerites._
_Spencerites insignis_ (Williamson). Fig. 192.
1878. _Lepidostrobus sp_., Williamson, Phil. Trans. R. Soc., p.
340, Pl. XXII.
1880. _Lepidostrobus insignis_, Williamson, Phil. Trans. R. Soc.,
p. 502, Pl. _XV_. figs. 8–12.
1889. _Lepidodendron Spenceri_, Williamson, Phil. Trans. R. Soc.,
p. 199, Pl. VII. figs. 20–22; Pl. VIII. fig. 19.
1897. _Spencerites insignis_, Scott, Phil. Trans. p. 83, Pls.
XII–XV.
Another type of lycopodiaceous strobilus, differing sufficiently from
_Lepidostrobus_ to deserve a special generic designation, is that
originally described by Williamson[439], from the Lower Coal-Measures
of Yorkshire, as a type of _Lepidostrobus_, _L. insignis_, but
afterwards[440] more fully investigated and assigned to a new genus
by Scott[441]. It should be pointed out that in a later publication
Williamson spoke of the lycopodiaceous axis, which he suspected might
belong to his _L. insignis_, as possibly worthy of recognition as a
distinct generic type.
[Illustration: FIG. 192. _Spencerites insignis_ (Williamson). (After
Miss Berridge.)]
Of the two species included by Scott in his genus _Spencerites_ only
one, _S. insignis_, need be considered. Since the publication of
Scott’s paper our knowledge of this type has been extended by Miss
Berridge[442] and by Prof. Lang[443].
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