Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
Still older stems of _Sigillaria_ are not uncommon from which the
leaf-scars and other superficial tissues have been exfoliated,
leaving exposed a longitudinally fissured surface of secondary cortex
characterised by pairs of considerably enlarged parichnos strands
(fig. 198) which are sometimes partially or wholly fused into one
(_Syringodendron_ state of _Sigillaria_). The single or double nature
of the elliptical or circular parichnos areas is doubtless due to the
degree of exfoliation, which may extend sufficiently deep into the
cortex to reach the level of the parichnos before the single strand has
bifurcated (cf. _Lepidodendron_, p. 100). In the Museums of Manchester,
Newcastle, and other places casts of large Sigillaria stems may be
seen, which illustrate the differences in breadth and regularity of
the vertical ribs, and in the size and shape of the parichnos areas
in different regions of a partially decorticated stem. A cast of a
ribbed species in the Manchester Museum, having a length of 185 cm.
and a breadth of 56 cm., shows in the upper portion straight vertical
grooves and broad ribs bearing pairs of parichnos scars 11 mm. long;
in the lower portion the ribs tend to become obliterated and the
parichnos scars, 2 cm. in length, may be partially fused and arranged
in much less regular vertical series. A feature of these older ribbed
Sigillarian stems is the increase in the number of the ribs from below
upwards. Kidston[476] has described a specimen in the Sunderland
Museum, 6 feet 6 inches long, with a circumference at the slightly
bottle-shaped base of 5 feet. On the lower portion of the stem there
are 29 broad ribs; about one-third the height many of these bifurcate,
producing as many as 40 ribs in the upper part where the cast has a
circumference of 3 feet. The increase in number of the ribs is due in
part to bifurcation, but also to the intercalation of new ones. As
Kidston points out, this example shows that as a stem grew in length
additional leaves were developed at the apex. A similar stem, which
illustrates very clearly the increase in the number of ribs from below
upwards, may be seen in the Newcastle Museum.
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