Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
Dr Kidston[496] is of opinion that certain species of _Sigillaria_
bore cones sessile on large vegetative shoots characterised by two
opposite rows of cup-like depressions like those in the Ulodendron
form of _Lepidodendron Veltheimianum_ (fig. 157). He has described
the Ulodendron condition of two species, _Sigillaria discophora_
(König) and _S. Taylori_ (Carr.); the cup-like depressions may have a
diameter of several centimetres and are distinguished from those of
_Bothrodendron_ by the almost central position of the umbilicus. The
specimens which he figures as _S. discophora_ are identified by him
with the stem figured by König as _Lepidodendron discophorum_ and by
Lindley and Hutton[497] as _Ulodendron minus_. We have already dealt
with the nature of Ulodendron shoots, expressing the opinion that in
spite of the often quoted specimen described by D’Arcy Thompson[498],
in which a supposed cone occurs in one of the cups, there is no
satisfactory case of any undoubted cone having been found attached to
the large Ulodendron scars. It is more probable that the Ulodendron
depressions represent the scars of branches, either elongated axes,
or possibly in some cases deciduous tuberous shoots which served as
organs of vegetative reproduction. A specimen figured by Kidston as
_Sigillaria Taylori_ from the Calciferous sandstone of Scotland[499]
bears a row of slightly projecting “appendicular organs” attached to a
Ulodendron axis; but these furnish no proof of their strobiloid nature.
The main question is, are these Ulodendron shoots correctly identified
by Kidston as Sigillarian? The surface of the specimens shows crowded
rhomboidal scars surrounded in some cases by a very narrow border
or cushion; the general appearance is, as Kidston maintains, like
that of _Sigillaria Brardi_ in which the leaf-scars are contiguous
(e.g. fig. 203, upper part). None of the leaf-scars exhibit the three
characteristic features, the leaf-trace and parichnos scars, but only
one small scar appears on each leaf-base area. In a more recent paper
Kidston figures a small piece of a stem from Kilmarnock, which he
identifies as _Sigillaria discophora_, showing the three characteristic
scars on the leaf-base area. There is no doubt as to the Sigillarian
nature of this specimen, but it is not clear if the piece figured is
part of a Ulodendron shoot[500].
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