Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The genus _Dipteris_, formerly included in the Polypodiaceae, has been
assigned to a separate family partly on account of the slight obliquity
of the vertical annulus (fig. 231, G) and on other grounds[702]. The
four species _Dipteris conjugata_, _D. Wallichii_, _D. Lobbiana_ (=
_D. bifurcata_), and D. _quinquefurcata_ (fig. 231) are characterised
by a creeping rhizome bearing fronds reaching a length of 50 cm.; in
_D. conjugata_ and _D. Wallichii_ the lamina is divided by a median
sinus into two symmetrical halves, while in other species the leaf
is dissected into narrow linear segments. The main dichotomously
branched ribs are connected by lateral branches and these by tertiary
veins, the delicate branches of which end freely within the square or
polygonal areolae (fig. 231, A′, E). The naked sori are composed of
numerous sporangia and filamentous hairs: while in some species the
soral development conforms to that characteristic of the Mixtae, it has
been shown that in one species, _D. Lobbiana_ (= _D. bifurcata_[703]),
the sporangia develope simultaneously as in the Simplices. _Dipteris_
occurs in company with _Matonia_ on Mt Ophir and elsewhere in the Malay
peninsula; it extends to the Philippines, Samoa, New Caledonia, China,
New Guinea, and the subtropical regions of Northern India.
• • • • •
The impossibility of drawing a hard and fast line between the divisions
adopted in any system of classification is well illustrated by the
ferns. In the main, the three-fold grouping suggested by Bower is
probably consistent with the order of evolution of the true ferns.
The Polypodiaceae, which are now the dominant group, are in all
probability of comparatively recent origin, while the Gradatae and
Simplices represent smaller subdivisions with representatives in remote
geological epochs. The genera _Loxsoma_, _Matonia_ and _Dipteris_
afford examples of ferns exhibiting points of contact with more
than one of Bower’s subdivisions: they are generalised types which,
like many relics of the past, are now characterised by a restricted
geographical range.
[Sidenote: RECENT FERNS]
[Illustration: FIG. 232. _Davallia aculeata._ (⅖ nat. size.)]
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