Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
[Illustration: FIG. 253. _Osmundites Kolbei_ Sew. (⅓ nat. size.)]
In another Jurassic species, _Osmundites Gibbiana_[787], the xylem is
of the _Osmunda_ type and consists of about 20 strands instead of a
continuous or almost continuous cylinder.
[Illustration: FIG. 254. _Osmundites Kolbei._ (Leaf-scars.)]
_Osmundites Kolbei_ Seward, figs. 253–255.
This species was founded on a specimen obtained by Mr Kolbe from
the Uitenhage series of Cape Colony[788]. The fossil flora and fauna
of this series point to its correlation with the Wealden or Neocomian
strata of Europe[789]. The type-specimen consists of several pieces
of a stem (fig. 253) which reached a length of about 90 cm. On the
weathered surface the remains of petiole-bases are clearly seen and
on the reverse side of the smaller piece shown in the figure numerous
sinuous roots are present in association with the leaf-stalks. The
depression _c_ in the larger specimen may mark the position of a
branch: at _a_ fig. 253 (enlarged in fig. 254, _a_) the vascular strand
of a petiole is exposed as a broad =U=-shaped band and at _b_ (fig.
254, _b_) the form of the petiole-bases is clearly shown[790]. With the
stem were found imperfectly preserved impressions of fronds referred to
_Cladophlebis denticulata_, a common type of leaf which was found also
in association with the slightly older New Zealand stem, _Osmundites
Dunlopi_.
[Illustration: FIG. 255. _Osmundites Kolbei_ Sew. Transverse section,
from a photograph supplied by Dr Kidston and Mr Gwynne-Vaughan.
(2½ nat. size.)]
An examination of the internal structure of the South African stem
by Dr Kidston and Mr Gwynne-Vaughan has revealed many interesting
features, which will be fully described in Part IV. of their Monograph
on fossil Osmundaceous stems. I am greatly indebted to these authors
for allowing me to publish the following note contributed by Dr
Kidston:—
“The section of _Osmundites Kolbei_ Seward, shown in fig. 255, presents
the usual appearance of an Osmundaceous stock. The parts contained in
this section are the stele, inner and outer cortex and a portion of
the surrounding mantle of concrescent leaf-bases. The whole specimen
has suffered much from pressure, but if restored to its original form
the xylem ring must have been about 19 mm. in diameter. The number of
xylem strands is about fifty-six and several of them are more or less
joined as in the modern genus _Todea_. The tracheae are of the typical
Osmundaceous type, that is to say, the pits are actual perforations and
several series of them occur on each wall of the larger tracheae.
Public-domain text, read in full here on John Shaqi.
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