Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
1851. _Sphenopteris nephrocarpa_, Bunbury, Quart. Journ. Geol. Soc.
Vol. VII. p. 129, Pl. XII. fig. 1.
1876. _Thyrsopteris Murrayana_, Heer, Flor. Foss. Arct. Vol. IV. (2)
p. 30, Pls. I. II. VIII.
The above list represents a small selection of the names applied to
Jurassic ferns from different localities which there are good grounds
for regarding as referable to a single type[904].
Frond tripinnate; pinnae linear acuminate, attached to the rachis
at a wide angle; the pinnules vary considerably in size and
shape; in some the lamina is divided into a few broad and rounded
lobes (fig. 275, B) while in others the leaflets are dissected
into narrow linear segments. The sori are borne at the ends of
veins; the fertile pinnules have a much reduced lamina and, in
extreme cases, bear a close resemblance to those of _Thyrsopteris
elegans_ (fig. 229, A, p. 294). The sori are partially enclosed in
a cup-like indusium and the sporangia appear to have an oblique
annulus.
Venation and habit of frond of the _Sphenopteris_ type.
[Illustration: FIG. 271. _Coniopteris hymenophylloides_ (Brongn.).
Nat. size. From a specimen in the Manchester Museum.]
The pinna shown in fig. 271 is the type-specimen of _Sphenopteris
arguta_ Lind. and Hutt. from the Yorkshire Inferior Oolite and is
indistinguishable from the English examples on which Brongniart founded
his species _S. hymenophylloides_. Fig. 272 shows a specimen from
the York Museum illustrating the difference between the sterile and
fertile pinnae. The resemblance of some fertile pinnae of _Coniopteris
hymenophylloides_ to those of _Thyrsopteris elegans_ has led to a
frequent use, without any solid justification, of the generic name of
the Juan Fernandez fern for Jurassic and Wealden plants. It is not
impossible that some of the fossils described by Heer from Jurassic
rocks of Siberia[905] as species of _Thyrsopteris_ are Cyatheaceous
ferns, but it is impossible to say with certainty that they are
generically identical with the recent species. In his monograph of the
Potomac flora of Virginia[906] and Maryland, Fontaine has described as
species of _Thyrsopteris_ several specimens of fronds which afford
no evidence as to the nature of the sori or sporangia. Some of the
fronds referred by this author to _Thyrsopteris rarinervis_[907],
which I examined in the Washington Museum, are in all probability
examples of _Onychiopsis_, a genus included in the Polypodiaceae. The
fragments described by Lester Ward[908] as species of _Thyrsopteris_
from the Lower Cretaceous of the Black Hills of North America afford
no satisfactory evidence of relationship to the recent type. Similarly
Velenovský has described a Lower Cretaceous _Onychiopsis_ from
Bohemia[909] as a species of _Thyrsopteris_, although the fertile
segments bear little or no resemblance to those of the Cyatheaceous
genus. Some fertile portions of fronds described by Heer[910] as
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