Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
This generic name, proposed by the late Professor Weiss[988],
is applied to a type of fructification illustrated by the plant
which Brongniart named _Pecopteris unita_, a species common in the
Upper Coal-Measures of England[989]. It is adopted by Kidston for
fertile specimens from Radstock which he describes as _Ptychocarpus
oblongus_[990], but the precise nature of the fertile pinnules of this
species cannot be determined.
_Ptychocarpus unita_ (Brongn.[991]). Fig. 291, A, B. (= _Goniopteris
unita_, Grand’Eury.)
This species has tripinnate fronds with linear pinnae bearing
contiguous pinnules of the _Pecopteris_ type (fig. 291, B), 4–5 mm.
long, confluent at the base or for the greater part of their length.
On the under surface of the fertile segments, which are identical
with the sterile, occur circular synangia (fig. 291, A) consisting
of seven sporangia embedded in a common parenchymatous tissue and
radially disposed round a receptacle supplied with vascular tissue.
The synangium is described as shortly stalked like those of _Marattia
Kaulfussii_ (fig. 245, B′, p. 320). In shape, in the complete union of
the sporangia, and presumably in the apical dehiscence, _Ptychocarpus_
agrees very closely with _Kaulfussia_ (fig. 245); but we cannot be
certain that we have not a collection of microsporangia simulating a
fern synangium.
A synangium closely resembling _Ptychocarpus_ has been described
by Mr Watson[992] from the Lower Coal-Measures of Lancashire as
_Cyathotrachus altus_, but there is no convincing evidence as to the
nature of the plant on which it was borne.
_Danaeites._
This generic name, instituted by Goeppert[993], has been used by
authors without due regard to the nature of the evidence of affinity to
_Danaea_. The type named by Stur _Danaeites sarepontanus_[994] (fig.
291, E) bears small pecopteroid pinnules with ovoid sporangia in groups
of 8–16 in two contiguous series on the lower face of the lamina. The
sporangia dehisce by an apical pore and are more or less embedded in
the mesophyll of the segments. No figures have been published showing
any detailed sporangial structure, and such evidence as we have is
insufficient to warrant the conclusion that the resemblance to _Danaea_
is more than an analogy.
_Parapecopteris._
_Parapecopteris neuropteroides_, Grand’Eury. Fig. 290, D.
The plant described by Grand’Eury[995] from the Coal-fields of Gard
and St Étienne, and made the type of a new genus, is characterised
by pinnules intermediate between those of _Pecopteris_ and
_Neuropteris_[996] and by the presence of two rows of united sporangia
along the lateral veins, as in _Danaea_ and _Danaeites_.
_Asterotheca._
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