Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The Upper Carboniferous fronds of a delicate Sphenopteris habit, to
which this name was assigned by Kidston[1023], were described by
him as _Eusphenopteris tenella_ (Brongn.)[1024] and compared with
Hymenophyllaceae; subsequently Kidston expressed the opinion that
_Urnatopteris_ may be a Marattiaceous fern, as Williamson[1025]
believed; he has since suggested that the sporangia are the
microsporangia of a Pteridosperm[1026]. The sterile and fertile pinnae
differ in the absence of a lamina in the latter. The sporangia (or
microsporangia) are characterised by a poricidal dehiscence.
The records from strata higher in the geological series than the
Permian, disregarding many of doubtful value, afford ample testimony to
the existence of Marattiaceae in Upper Triassic and Rhaetic floras.
_Marattiopsis._
The generic name _Danaeopsis_ was applied by Heer[1027] to an Upper
Triassic fern, previously described by Presl as _Taeniopteris
marantacea_. A splendid specimen from the Keuper of Stuttgart is
figured in Schimper’s Atlas[1028] showing the pinnate habit of the
frond and the broadly linear segments, 25 cm. × 3·5 cm., bearing rows
of contiguous sporangia. The large pinnules have a strong midrib
giving off curved and forked lateral veins. Presl’s species may most
appropriately be included in the genus _Marattiopsis_. A specimen of
_M. marantacea_ described by Leuthardt[1029] as _Danaeopsis marantacea_
from the Upper Trias of Basel shows a peculiarity in the venation;
the lateral veins often fork near their origin, as noticed by other
authors, but each vein forks a second time near the edge of the lamina
and the two arms converge, forming a series of intramarginal loops
(fig. 265, B).
_Marattiopsis Muensteri_ (Goepp.). Fig. 245, D, E.
This widely spread Rhaetic plant affords the best example of a
post-Permian species which may be accepted as an authentic record of
fossil Marattiaceae. Various generic names have been used for this
species; Goeppert originally described the plant as _Taeniopteris
Muensteri_[1030]; Schimper[1031] proposed the name _Marattiopsis_, and
Schenk[1032] substituted _Angiopteris_ on the ground that the fertile
pinnules resemble that genus rather than _Marattia_. _Marattiopsis_, if
interpreted as indicating a _family_ resemblance rather than special
affinity to the genus _Marattia_, would seem to be the more appropriate
designation.
This species has been figured by several authors and in many instances
with fertile pinnules; the best illustrations are those published by
Zeiller[1033] in his monograph of Tonkin plants.
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