Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
Phloem elements, recognisable by the presence of a thickened reticulum
enclosing small sieve-areas (fig 133, B, _s_) are fairly abundant,
and for the rest this intracambial region is composed of thin-walled
parenchyma. In longitudinal section these tissues present an appearance
almost identical with that observed in a transverse section. Fig.
133, B represents a longitudinal section, through the intracambial
zone and the edge of the stele, of a younger stem than that shown
in fig. 133, A. Most of the radially disposed cells internal to
the meristematic region are parenchymatous without any distinctive
features; a few scattered sieve-tubes (_s_) are recognised by their
elliptical sieve-areas and an occasional tracheid can be detected. The
cambium cuts off externally a succession of segments which constitute
additional cortical tissue (fig. 133, A, _cr_) of homogeneous
structure, composed of parenchymatous cells containing starch and
rich in intercellular spaces. As the stem grows in thickness the
secondary cortex reaches a considerable breadth and the superficial
layers are from time to time exfoliated as strips of dead and crushed
tissue (fig. 133, A, _b_). The diagrammatic sketch reproduced in fig.
133, A, serves to illustrate the arrangement and relative size of the
tissue-regions in an _Isoetes_ stem. In the centre occur numerous
spirally or reticulate tracheae scattered in parenchymatous tissue
which has been considerably stretched and torn in the peripheral
region of the stele; the radiating lines mark the position of the
leaf-traces (_lt_) in the more horizontal part of their course. The
zone between the cambium (_c_) and the edge of the central cylinder
consists of radially disposed secondary tissue of short, and for the
most part unlignified, elements including sieve-tubes and parenchyma;
the secondary xylem elements consist largely of thin-walled rectangular
cells with delicate spiral bands, but discontinuous rows of lignified
tracheae (_x²_) occur in certain regions of the intracambial zone. The
rest of the stem consists of secondary cortex (_cr_) with patches of
dead tissue (_b_) still adhering to the irregularly furrowed surface.
The structure of the cambium and its products is shown in the detailed
drawing reproduced in fig. 133, D. Many of the elements cut off on the
inner side of the cambium exhibit the characters of tracheids: most
of these are unlignified, but others have thicker and lignified walls
(_tr_).
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