Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
This generic name has been instituted by Zeiller[224] for specimens
from the coal basis of Blanzy (France). It is applied to heterosporous
species with the habit of _Selaginella_: Zeiller preferred the
designation _Selaginellites_ to _Selaginella_ on the ground that
the type species differs from recent forms in having more than four
megaspores in each megasporangium. It is, however, convenient to extend
the term to all heterosporous fossil species irrespective of the
spore-output.
_Selaginellites Suissei_ Zeiller.
This species was described in Zeiller’s preliminary note[225] as
_Lycopodites Suissei_, but he afterwards transferred it to the genus
_Selaginellites_. In habit the plant bears a close resemblance
to _Lycopodites macrophyllus_ of Goldenberg; the shoots, 1–3 mm.
thick, are branched in a more or less dichotomous fashion and bear
tetrastichous leaves. The larger leaves reach a length of 4–6 mm.
and a breadth of 2–3 mm.; the smaller leaves are described as almost
invisible, closely applied to the axis, oval-lanceolate and 1–2 mm.
long with a breadth of 0·5–0·75 mm. Long and narrow strobili (15
cm. by 8–10 mm.) terminate the fertile branches; these bear crowded
sporophylls with a triangular lamina and finely denticulate margin.
Oval sporangia were found on the lower sporophylls containing 16–24
spherical megaspores 0·6–0·65 mm. in diameter. The outer membrane
of the spore is characterised by fine anastomosing ridges and thin
plates radiating from the apex and forming an equatorial collarette.
The microspores have a diameter of 40–60μ and the same type of outer
membrane as in the megaspores. The megaspores of the recent species
_Selaginella caulescens_, as figured by Bennie and Kidston[226],
resemble those of the Palaeozoic type in the presence of an equatorial
flange. It is interesting to find that, in spite of the occurrence of
16–24 megaspores in a single sporangium the size of the fossil spores
exceeds that of the recent species.
_Selaginellites primaevus_ (Gold.). Fig. 135, A, fig. 138.
1855 _Lycopodites primaevus_, Goldenberg[227].
1870 _Lycopodium primaevum_, Schimper[228].
1907 _Selaginellites primaevus_, Halle[229].
[Illustration: FIG. 138. _Selaginellites primaevus_ (Gold.). (After
Goldenberg.)]
In habit this species, first recorded by Goldenberg from the
Coal-Measures of Saarbrücken, is similar to _S. Suissei_ Zeill.
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