Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
A dual system of terminology has been unavoidably adopted for
species of _Lepidodendron_: the majority of specific names have been
assigned to fossils known only in the form of casts or impressions,
while petrified fragments, which unfortunately seldom show the
surface-features, have received another set of names. A glance at the
older palaeobotanical literature reveals the existence of several
generic designations, which fuller information has shown to have been
applied to lepidodendroid shoots deprived of some of their superficial
tissues before fossilisation and differing considerably in appearance
from the more complete branches of the same species[240]. It has in
some instances been possible to correlate the two sets of specimens,
casts or impressions, showing external features, and petrified
fragments. We may reasonably expect that future discoveries will enable
us to piece together as definite specific types specimens at present
labelled with different names.
A well-preserved leaf-cushion of a _Lepidodendron_—the most obvious
distinguishing feature of the genus—is rhomboidal or fusiform and
vertically elongated (fig. 146, C, E; fig. 185, C, D): in exceptional
cases it may reach a length of 8 cm. and a breadth of 2 cm. The
cushion as a whole represents a prominent portion of the stem or
branch comparable with the elevation on the twig of a Spruce Fir and
the leaf-base of a _Lycopodium_ (cf. fig. 121, A, lower portion) which
appears in a transverse section of a branch as a rounded prominence
(cf. _Lycopodium_, fig. 125, A and H). Disregarding differences in
detail, a typical Lepidodendron leaf-cushion is characterised by a
clearly defined smooth area often situated in the middle region (fig.
146, C, _s_). This is the leaf-scar or place of attachment of the base
of the leaf which was cut off by an absciss-layer while the branch was
comparatively young, as in recent forest trees and in some species of
Ferns. On the leaf-scar are three smaller scars or cicatricules, the
central one is circular or more or less triangular in outline, the two
lateral scars being usually oval or circular. The central pit marks
the position of the single vascular bundle which constituted the
conducting tissue connecting the leaf with the main vascular system of
the stem. The two lateral scars (figs. 145, A, _p_; 146, C, _s_; 147,
_p_) represent the exposed ends of two strands of tissue, the forked
branches of a strand which pass from the middle cortex of the stem into
the leaf; this is known as the parichnos, a name proposed by Professor
Bertrand in 1891[241].
The specimen shown in fig. 141 shows the linear leaves attached to
their respective cushions.
[Illustration: FIG. 141. _Lepidodendron Sternbergii._ From a specimen
in the British Museum (No. v. 1235) from the Coal-Measures of
Shropshire. (Nat. size.)]
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