Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The area of the cushion excluding the leaf-scar is spoken of by some
writers as the field. Below the leaf-scar the kite-shaped cushion
tapers to a gradually narrowing basal position: in _Lepidodendron
Veltheimianum_, a species characteristic of Lower Carboniferous
strata, it is seen to be continuous, as a ridge with sloping sides,
with a lower cushion (fig. 185).
Below a leaf-scar the cushion frequently shows a pair of oval areas on
which a fine pitting may be detected in well-preserved impressions,
these oval scars, as seen in fig. 185, D, are practically continuous
at the upper end with the parichnos scars on the leaf-scar area;
this is explained by the fact that these infra-foliar scars also owe
their existence to patches of lacunar, aerenchymatous tissue in close
connexion with the parichnos[243].
Shortly before entering the base of the leaf-lamina the parichnos
divides into two arms which diverge in the outer cortical region right
and left of the vascular bundle, and passing obliquely upwards they
come close to the surface of the leaf-cushion just below the leaf-scar.
The diagram—fig. 144, B—shows a leaf-trace, _lt_, in the leaf-cushion,
as seen in a diagrammatic drawing of a vertical radial section of a
stem, the dotted lines, _p_, _p′_, show the two parichnos arms which
are represented as impinging on the surface of the leaf-cushion at
_p′_, and then bending upwards to pass into the leaf-base right and
left of the vascular bundle or leaf-trace. For convenience the arms of
the parichnos are represented in one plane though actually in different
vertical planes.
Fig. 144, A, shows the difference between a view of the original
surface of a _Lepidodendron_, as at _a_, where a leaf-cushion with a
leaf-scar is seen, and a view of an impression representing the outer
cortex, _b_, a short distance below the surface. The surface _b_,
in fig. 144, A, corresponds to the face _d_-_e_ in the diagrammatic
longitudinal section fig. 144, B: the outline of each cushion is
clearly visible and in the centre is seen the leaf-trace, _lt_, with
its parichnos.
The surface-features, _a_ (fig. 144, A), have been impressed on the
rock, _c_, (fig. 144, B) in which the specimen was entombed and by the
removal of the cast of the stem, that is the thickness _b_ to _e_ in
fig. 144, B, the form of the leaf-cushion is revealed. The presence of
the two infra-foliar parichnos scars at _p′_ (fig. 144, A) is explained
by the diagram, fig. 144, B, _p′_.
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