Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The drawing reproduced in fig. 146, A, showing the leaf-scar on the
upper edge of the cushion should have been reversed with the leaf-scars
pointing downwards. This figure represents part of the surface of a
specimen consisting of the outer cortex of a stem with leaf-cushions 3
cm. broad. The thickness of this specimen is 4 cm.: a section through
the line _ab_ is represented in fig. 146, D (reproduced in the correct
position, with the leaf-scars, _sc_, pointing downwards): internal
to the cushions is a band of secondary cortex (the shaded strip on
the outer edge of the section) which was formed on the outside of
the phellogen. The phellogen is a cylinder of actively dividing cells
in the outer part of the cortex of the stem, often spoken of as the
cork-cambium or cortical meristem, which produces a considerable amount
of secondary cortical tissue on its inner face and a much smaller
amount towards the stem surface. This delicate cylinder frequently
forms a natural line of separation between the outer shell of bark and
the rest of the stem. In the specimen before us, the thin-walled cells
of the phellogen were ruptured before petrification and the outer shell
of bark was thus separated as a hollow cylinder from the rest of the
stem: this cylinder was then flattened, the two inner surfaces coming
into contact. Fig. 146, D, represents a section of one half of the
thickness of the flattened shell.
This separation of the outer cortex, and its preservation apart
from the rest of the stem, is of frequent occurrence in fossil
lycopodiaceous stems. The flattened outer cortical shell of a
_Lepidophloios_, specifically identical with that shown in fig. 146, A
and D, was erroneously described by Dr C. E. Weiss in 1881 as a large
lepidodendroid cone[258].
Fig. 146, B, affords a view of the inner face of the specimen of which
the outer surface is seen in fig. 146, A: the surface shown in the
lower part of the drawing, on which the boundaries of the cushions are
represented by a reticulum, corresponds to the inner edge of the strip
of secondary cortical tissue represented by the vertically shaded band
in fig. 146, D.
The shaded surface in fig. 146, B, represents a slightly deeper level
in the stem which corresponds to the outer edge of the vertically
shaded band of fig. 146, D: the narrow tapered ridges (fig. 146, B)
represent the leaf-traces passing through the secondary cortex, and the
fine vertical shading indicates the elongated elements of which this
strip of secondary cortex is composed.
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