Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The inner edge of the phelloderm seen at _e_ in fig. 151, C, passes
suddenly into the inner primary region of the outer cortex (fig. 151, A
and C, _g_) which comprises two types of parenchymatous tissue, patches
of isodiametric cells, _g_, _g_, alternating with radially arranged
areas consisting of tangentially elongated elements (fig. C, _f_, _f_;
fig. D) which extend as wedges into the phelloderm.
The longitudinal section represented in fig. 152, B, shows an equal
bifurcation of a stem in which no secondary xylem is present; in the
lower part of the section the xylem and the outgoing leaf-traces are
seen in radial section and at the upper end of each arm the leaf-traces
alone, _lt_, are exposed, as in fig. 148, B. It is interesting to
notice the large amount of phelloderm which has been produced in the
fork of the branch, at _pd_, where greater strength is required.
[Illustration: FIG. 152. _Lepidodendron vasculare._ Sections of
dichotomously branched shoot.
A. From a section (10·5 × 9 cm.) in the Cambridge Botany School
Collection.
B. From a section (8 cm. long) in the Cambridge Collection.]
The section represented diagrammatically in fig. 152, A, has lost the
outermost part of the cortex together with the leaf-cushions; it
consists largely of secondary cortex composed of radially disposed
phelloderm cells and tangentially placed secretory strands (represented
by the discontinuous black lines in the drawing): the dotted region
in the central part of the axis is composed of primary cortical
parenchyma, and the two spaces surrounding the steles contain portions
of the lacunar middle cortex. Each stele possesses a narrow crescentic
zone of secondary xylem; the amount is greater in the case of the
right-hand stele, of which a small piece is shown on a larger scale;
the striking contrast in size between the outer and more internal
secondary tracheae is no doubt the expression of some unfavourable
condition of growth. The position of the secretory zone beyond the
secondary xylem is shown at _sc_, fig. 152, A.
[Illustration: FIG. 153. _Lepidodendron vasculare._
(From a specimen (16 × 7·5 cm.) in the Manchester Museum.)]
An example of a large and partially decorticated stem is afforded by
the specimen (16 × 7·5 cm.) shown in fig. 153. The irregularly ribbed
surface is formed of rather thick-walled phelloderm, in which occur
tangentially arranged rows of secretory strands. The tapered form of
the secondary cortex as it abuts internally on the primary cortex is
shown very clearly in the drawing (cf. fig. 151, C). The stele in this
much older stem consists mainly of secondary wood.
[Illustration: FIG. 154. _Lepidodendron vasculare._ Shoot (2·8 cm.
diam.) with two steles.
(From a specimen from Halifax in the Williamson Collection, British
Museum, No. 340.)]
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