Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The _Knorria_ state represents the impression or cast of the outer
cortical region too deep below the leaf-cushion region to retain
any indications of the cushion-form; the ridges are the casts of the
spaces produced in the cortex by the decay of the sheath of delicate
cells surrounding each leaf-trace and by the decay of the thin-walled
cells of the parichnos. The occasional forked apex of a ridge is the
expression of the fact that the cast was made at the region where the
parichnos divides into two arms (cf. p. 100). In certain specimens it
is possible to connect the Knorria casts with associated lepidodendroid
stems which may be determined specifically; but when we have no
evidence as to surface-features the fossils may be designated casts
of lepidodendroid stems in the Knorria condition. Such casts are
illustrated by numerous drawings in palaeobotanical literature[277].
_b._ _Bergeria._
This is another name first used by Sternberg in his classic work,
_Die Flora der Vorwelt_, for casts of lepidodendroid plants such as
Steinhauer[278] had previously figured as _Phytolithus cancellatus_.
Brongniart[279] recognised that the application of the generic title
_Lepidodendron_ should be extended to include specimens referred by
Sternberg to _Bergeria_, and a few years later Goldenberg[280] realised
that this name does not stand for well-defined generic characters.
The correctness of these views was, however, first satisfactorily
demonstrated by Carruthers[281] and by Feistmantel[282].
If a Lepidodendron stem loses its superficial layers of outer
cortex and in this condition is embedded in sand or mud, the cast
is distinguished from that of a perfect stem by the absence of
the leaf-scars and by other features. It may, however, still show
spirally disposed areas, corresponding approximately to the original
leaf-cushions, which are characterised by a small depression or pit
either at the apex or near the centre of each oval area: the pit marks
the position of the leaf-trace and its parichnos strand. In some
cases the exposed surface may be smooth without any indication of
leaf-cushions, while narrow spirally arranged grooves represent the
obliquely ascending vascular bundles passing through the cortex to the
leaves.
Fig. 185, B, shows the Bergeria state of _Lepidodendron Veltheimianum_,
which differs from the Knorria condition in the fact that decortication
had not extended below the level at which the form of the leaf-cushions
could be recognised. It is clear that no sharp line can be drawn in all
cases between the different degrees of decortication as expressed by
the terms _Knorria_ and _Bergeria_.
A list of synonyms of _Knorria_, _Bergeria_, and _Aspidiaria_ forms of
stem and a detailed treatment of their characteristic features may be
found in a recent work by Potonié[283].
_c._ _Aspidiaria._
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