Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
This species[541] affords a good illustration of the generic
characters already summarised. The strap-like leaves are fleshy
and the occasionally forked, parallel or slightly divergent, veins
are embedded in a homogeneous mesophyll with hypodermal strands of
mechanical tissue. The pith consists of parenchyma in vertical series
with scattered secretory sacs and differs from that of _Cordaites_
and _Mesoxylon_ in the absence of transverse discs. There are 13
primary-xylem strands close to the inner edge of the secondary wood:
the centripetal tracheids are scalariform or have multiseriate pitting
like that in the secondary xylem. The structure of the leaf-traces is
clearly shown in fig. 464: the double trace seen in fig. 464, C, has
two protoxylem-strands accompanied by some parenchyma, and these are
almost enveloped by the metaxylem tracheids which abut on the secondary
wood. At this stage in its course, that is just before bending
outwards, the centripetal xylem reaches its maximum development and the
trace forms a prominent and broad twin-strand in striking contrast to
the two narrower and tangentially extended strands shown in fig. 464,
E, D. Each of these strands with a single protoxylem-group would at a
higher level assume the broader and more compact form and contain two
protoxylems as in fig. 464, C. The tracheids of the secondary xylem
have 4–7 alternate rows of contiguous alternate pits on the radial
walls: the medullary rays are 2–3 cells broad and may be 60 cells deep.
According to Renault[542] several small oblique pits occur on the
radial walls of the ray cells. The secondary phloem, separated by a
normal cambium from the xylem, forms a broad band of sieve-tubes with
lateral sieve-plates like those in _Medullosa anglica_ alternating
with tangential rows of parenchyma. The cortex is relatively narrow
and in older stems is chiefly occupied by secondary tissue formed from
deep-seated phellogens.
[Illustration: Fig. 464. _Poroxylon_. A, B, _Poroxylon Boysseti_.
A, transverse section of leaf in the region of lateral expansion of
the petiole; B, portion of vascular tissue of A. C–E, _Poroxylon
Edwardsii_. C, leaf-trace showing recent separation of the two
protoxylems; D, leaf-trace at a lower level with more widely separated
protoxylems, _px_; E, leaf-trace intermediate between C and D. (From
photographs supplied by Prof. Bertrand.)]
_Poroxylon Boysseti_ Renault.
The stems of this species agree closely with those of _P. Edwardsii_,
the chief difference being in the structure of the secondary phloem
which does not show the regular concentric alternation of sieve-tubes
and parenchyma.
_Poroxylon stephanense_ Bertrand and Renault.
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Elsewhere in the archive
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account