Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
This widely distributed and mainly
Palaeozoic genus is especially well represented in the coalfields of
France where in some localities it contributed largely to the formation
of seams of coal[544], and it is chiefly from the researches of French
Palaeobotanists that our knowledge of its morphology is derived.
_Cordaites_ has shared the fate of most other abundant fossil plants
in the distribution of its _disjuncta membra_ among several genera
and classes, but on the whole the information that is now available
enables us to reconstruct the complete plant with a greater degree of
confidence than is usually attainable.
_Cordaites_ may be described as a forest-tree closely resembling
in habit and probably in size the recent Conifer _Agathis_, more
especially such species as _A. macrophyllus_, _A. vitiensis_ and others
with leaves considerably longer than those of the Kauri Pine (_A.
australis_)[545]. The main stem reached a considerable height before
giving off scattered branches bearing spirally disposed, sessile, and
often crowded leaves[546] like the foliage of _Agathis_. The absence of
any evidence of a two-ranked arrangement of leaves on lateral branches
suggests a general tendency towards a vertical rather than a horizontal
direction of growth. The sessile and closely set leaves for the most
part of leathery texture vary considerably in length and breadth in
different types (figs. 466–472): in some the broadly linear lamina with
its parallel veins and perfectly constructed I-shaped girders (fig.
465) reached a length of nearly 100 cm., in shape like the blade of
a straight broad-sword or the leaves of a _Yucca_, torn by the wind
into strips; in other forms the lamina is shorter and more obovate,
while in some the leafy shoots must have looked like slender stems of
the smaller-leaved Bamboos. There is no proof that young vegetative
branches with their spirally rolled leaves[547] were protected by
bud-scales, but some oval triangular scales (fig. 468, C), occasionally
found in association with larger foliage-leaves, may have served that
purpose. The branches from which leaves had recently fallen at the
time of fossilisation are characterised by transversely elongated
oval scars, occasionally showing a slightly curved row of pits like
the marks of leaf-traces on the scars of a Horse Chestnut, sometimes
terminating a feebly projecting decurrent leaf-cushion (fig. 466, C).
The leaves persisted for a comparatively long period as in _Araucaria
imbricata_, and on older leafless branches the scars are transversely
stretched; the leaf-cushion loses its individuality and eventually the
development of secondary cortical tissue causes the exfoliation of the
superficial bark.
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