Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
On the other hand the statement that annual rings
occur in Palaeozoic wood is often incorrect, partial rings having been
confused with regular concentric cylinders of summer elements. Dawson
and Matthew[668] described rings in _D. ouangondianum_, and Goeppert
and Stenzel[669], who examined the Canadian material, refer to circles
like annual rings; but Penhallow[670] states that there is no evidence
of true growth-rings.
The medullary rays are uniseriate and consist of thin parenchymatous
cells with unpitted walls; they vary considerably in depth, usually
comparatively shallow but in some cases 40 or 50 cells deep. In recent
Araucarineae the rays are generally shallower. The absence of special
receptacles, other than occasional resiniferous tracheids, for products
of secretion is a feature common to _Dadoxylon_ and the Araucarineae.
The phloem presents no features of special interest, but our knowledge
of this tissue is comparatively meagre.
Among other examples of large _Dadoxylon_ stems some of which no
doubt bore Cordaitean foliage—though as a rule we have insufficient
information as regards anatomical characters to enable a decision
to be made between _Cordaites_ and _Mesoxylon_—reference should be
made to the imposing array of silicified trunks in the grounds of the
Chemnitz Museum[671]. These were obtained from Lower Permian strata
at Hilbersdorf near Chemnitz from beds overlain by porphyry tuff and
resting on quartz porphyry, the volcanic material which furnished
the siliceous solutions. Several large pieces of wood were found in
association with stems of _Medullosa_ and _Psaronius_, leaves of
_Cordaites_, _Artisia_ pith-casts, and _Cardiocarpus_ seeds with
specimens of _Walchia_, _Gomphostrobus_ and other plants. Sterzel
describes a stem 16·5 met. long and 1·5 met. in diameter; on the main
trunk the branch-scars are scattered but on some branches there is
a tendency to a whorled arrangement. This and many other stems are
referred to _Araucarioxylon_ (or _Dadoxylon_) _saxonicum_, a species
first described by Reichenbach as _Megadendron saxonicum_. In one
specimen Sterzel states that the bordered pits are generally in 1–2
rows, though rarely in 3–4 rows, on the radial walls of the tracheids
which they do not completely cover: the medullary rays reach a depth of
between 20 and 30 cells. It is noteworthy that the stem 16·5 met. long
has a pith-cast of the _Tylodendron_ type.
_Dadoxylon_ (_Cordaites_) _Brandlingii_ (Lindley and Hutton).
1831. _Pinites Brandlingii_ Lindley and Hutton, Foss. Flor. Vol. +i.+
Pl. +i.+
1850. _Araucarites Brandlingii_ Goeppert, Foss. Conif. p. 232, Pls.
+xxxix.–xli.+
1890. _Cordaioxylon Brandlingii_ Schenk, in Schimper and Schenk, p.
853, fig. 408.
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