Fig. 481, A, B, _Cordaianthus Grand’Euryi_ Ren., shows a longitudinal section of the nucellus, 1·5 × ·7 mm., and part of the integument of an ovule at the time of pollination, which was probably aided by the secretion of a drop of mucilage as in the ovules of recent Conifers. The integument, separated by a broad space from the nucellus, is cut in the plane of the two vascular strands, _v_. From the centre of its broad upper surface the nucellus projects upwards as a beak, _b_, and this originally engaged with the micropylar canal formed by the integument: the lighter patch below the beak is the pollen-chamber (fig. 481, B, _pc_) containing two microspores, and two more, _p_, are seen above the nucellar beak. In another species described by Renault, _C. Lacattii_, the nucellus fills the space bounded by the integument. * * * * * (_b_) _Staminate inflorescences._ The male inflorescence, though smaller, is similar in habit to the ovulate shoot: the secondary branch consists of a short axis bearing crowded, spirally disposed, bracts, and the actual flowers are represented by single stamens or groups of 2–3 highly specialised microsporophylls. Each microsporophyll consists of a long filament with a central vascular strand bearing at its apex 3–4 long microsporangia (fig. 481, F, _m_) which open longitudinally as seen in fig. 482, A. The microsporangia are 2·5–3 mm. long covered by dark palisade cells and thin-walled parenchyma, shown as indistinct patches in the photograph. Some of the elliptical and comparatively large microspores are seen in fig. 482, B; the exine is finely punctate and inside are the remains of a few thin cells in which presumably spermatozoids were developed. The microspores shown in fig. 482, B, have a maximum length of 0·1 mm.: Renault describes some as 0·9 mm. long while others are much smaller. Fig. 481, F, is a transverse section of a staminate inflorescence showing near the centre five groups of microsporangia, each sporangium having the form of a curved incomplete dark band indicating that dehiscence has occurred. [Illustration: Fig. 482. _Cordaianthus._ Microsporangia, A, and Microspores, B. (University College Collection, London.)] =Mesoxylon.= Scott and Maslen.