Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
_Archaeopitys Eastmanii_ Scott and Jeffrey. The genus
_Archaeopitys_[770] has recently been instituted for a specimen of a
stem from the Lower Carboniferous rocks of Kentucky which differs from
_Pitys antiqua_, the type with which it is most closely allied, in the
position and behaviour of the primary vascular strands in the pith. The
type-specimen is a piece of stem 2·7 cm. in diameter including a solid
parenchymatous pith 5·5 mm. broad and a cylinder of secondary wood. The
wood is similar to that of _Cordaites_; the tracheids have 2–3 rows of
pits but for the most part the details of structure are not preserved;
the medullary rays are both uniseriate and multiseriate and not very
deep; the structure is more Cordaitean than Cycadean. A characteristic
feature is the occurrence of about 30 vascular bundles in the medullary
region: these mesarch strands, with the protoxylem near the centre,
are scattered through the pith and several lie on the outer edge in
contact with the secondary xylem or are partially embedded in the
xylem-cylinder. The examination of a series of sections demonstrated
the fusion of perimedullary with medullary strands and the occasional
union of the strands of both regions with one another. It appears that
the perimedullary strands are the leaf-traces while the strands deeper
in the pith are merely branches from the peripheral leaf-trace strands.
In _Pitys antiqua_ the medullary xylem strands are confined to
the outer zone of the pith and constitute the leaf-traces: in
_Archaeopitys_, on the other hand, the medullary strands are
scattered through the pith and the leaf-traces are restricted to the
circummedullary region in actual contact with the secondary xylem. In
the Devonian genus _Callixylon_ there are similar strands but they are
confined to the edge of the pith and are usually in contact with the
wood as are the circummedullary strands in _Archaeopitys_. The grouping
of the secondary xylem of _Callixylon_ into wedge-shaped masses at the
inner edge of the cylinder is a characteristic feature; this feature is
less definite in _Archaeopitys_ and absent from _Pitys_.
=Callixylon.= Zalessky.
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