Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
Brongniart[905] instituted this genus[906] on an incomplete transverse
section of a seed from Grand’ Croix, 8 mm. in diameter, which he called
_Eriotesta velutina_, characterised by a ribbed and probably octagonal
testa bearing numerous elongated hairs over the whole surface. The
material is, however, too meagre to throw any light on the important
features of the seed.
=Gaudrya.= Grand’Eury.
_Gaudrya trivalvis_ Grand’Eury.
The generic name _Gaudrya_[907] was proposed for two petrified seeds
from the Gard coal-field briefly described as _G. trivalvis_, the
type-species, and _G. lagenaria_. The testa of the former species
shows signs of splitting along three equidistant lines; it consists
of a sarcotesta enclosing an endotesta with six longitudinal ribs:
Grand’Eury speaks of lacunae between the endotesta and nucellus which
he regards as spaces in a tissue which made the seed buoyant and
facilitated dispersal by water. It is not clear whether the nucellus
and integument were originally connected or laterally free: the lacunae
may be the remains of an inner flesh and not spaces in a spongy tissue.
The genus is radiospermic and characterised by a long micropyle and
a broad pollen-chamber. On the available evidence it is difficult to
assign the specimens to their systematic position, but the genus is
probably a member of the Trigonocarpales.
III. +Cardiocarpales.+
Platyspermic seeds for the most part belonging to Cordaitalean plants
and agreeing in their plan of organisation, even more closely than
the seeds of the Trigonocarpales, with those of recent Cycads. The
nucellus is free laterally from the integument and there is a series of
vascular bundles in the nucellus. The testa consists of an outer flesh,
the sarcotesta, which may reach a considerable thickness, a shell
or sclerotesta, and in some genera an endotesta. In seeds preserved
as impressions the sarcotesta often gives them a winged appearance
(_Samaropsis_). The pollen-chamber is relatively small, resembling in
this respect the chamber in recent Cycads. A central prolongation of
the prothallus-tissue in the form of a blunt column or ‘tent-pole’
is a characteristic feature; the same feature occurs in seeds of the
Lagenostomales, but in the Cordaitales it resembles more nearly the
‘tent-pole,’ as it was called by Hirase, in recent Cycadean seeds and
in _Ginkgo_. The presence of two vascular strands in the sarcotesta
in the principal plane of the seeds is a character shared with the
seeds of _Cycas_: the position and course of these bundles are useful
characters for distinguishing different types within the group. The
microspores are multicellular.
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