Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
It is proposed to restrict the name _Rhabdocarpus_[947] to impressions
and casts of seeds of the type represented by _R. tunicatus_ as figured
by Berger[948] and reproduced in fig. 506, K, the term _Rhabdospermum_
being applied to seeds of similar form in which are shown certain
distinguishing anatomical features. _Rhabdocarpus tunicatus_ Berger
is a species founded on a specimen from the Coal Measures of Silesia
characterised by an outer carbonised testa prolonged apically as a
blunt snout and, as seen in fig. 506, K, covering an apparently ribbed
nucule, but the ‘ribs’ are due to the presence of fibrous strands and
are not ribs in the ordinary sense. The outer tissue shows numerous
longitudinal striations due presumably to the presence of fibrous
elements in the sarcotesta like those shown in the petrified seed
represented in fig. 501, C. The genus is defined by Berger as follows:
‘Semina ovata vel elliptico-oblonga secundum longitudinem parallele
nervosa vel tenuissime striata, putamine (interdum deficiente)
instructa.’ As used by Berger and many other authors _Rhabdocarpus_
includes a miscellaneous collection of seeds often differing widely
from the type-species. Many of the examples correctly referred to
Berger’s genus are platyspermic though a bilateral symmetry is by no
means always clear. Renault and Zeiller[949] in their definition of
_Rhabdocarpus_ include bilateral symmetry as a characteristic feature
and speak of the seeds as oblong or oval with a pointed or truncate
apex and a rounded base. Impressions of _Rhabdocarpus_ differ from
those of _Cardiocarpus_ or _Cordaicarpus_ in their more elongate
form, always longer than broad, and in the absence of a basal sinus.
The seeds found attached to _Neuropteris_ pinnae and, in external
features, agreeing with many specimens included in Berger’s genus, have
been transferred by P. Bertrand[950] and Arber[951] to a new genus
_Neurospermum_[952]. Arber[953] in his recent revision of British seeds
proposes to restrict the name _Rhabdocarpus_ to platyspermic seeds
having a ‘large unsymmetrical nucule enclosed in a large unsymmetrical
wing or sarcotesta,’ that is to forms symmetrical in one plane. In
this category he includes _Rhabdocarpus tunicatus_ Berg. (fig. 506,
K) and _R. subtunicatus_[954] Grand’Eury, but it is not clear on what
grounds Berger’s species is spoken of as symmetrical in only one
plane: in the species _R. Lilleanus_ Arb.[955] the symmetry is hardly
sufficiently well defined to rank as a generic character. In the case
of the _Neuropteris_ seeds the apical snout is slightly curved, thus
giving them an unsymmetrical appearance (cf. fig. 422, p. 114). The
Carboniferous and Permian seeds usually referred to _Rhabdocarpus_
are transferred by Arber to his genus _Platyspermum_[956], a
designation for which it is proposed to substitute Nathorst’s genus
_Holcospermum_[957]. In seeds preserved as more or less flattened
impressions it is practically impossible in many cases accurately to
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