Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The platyspermic seeds included in this genus agree in size and form
with impressions assigned to _Rhabdocarpus_ and as regards the main
features conform anatomically to _Cardiocarpus_; they were probably
borne on Cordaitean plants. Fig. 501, C, represents a longitudinal
section of the species _Rhabdospermum cyclocaryon_ described
by Brongniart as _Rhabdocarpus cyclocaryon_: the sarcotesta is
particularly well developed in the apical region; at the apex a portion
of the micropyle is seen at _m_ and near the nucellus are pieces of
the sclerotesta shown in black. The presence of anastomosing fibres
near the surface is a characteristic feature: these, as Bertrand points
out, do not form a hypodermal tissue in the strict sense as they may
be separated by some of the thin-walled parenchyma of the sarcotesta
from the epidermis. The sclerotesta is only partially preserved but the
inner portion forms a dark line enclosing the nucellus, the superficial
tissue of which is separated from the shrunken prothallus represented
by the almost spherical dotted region: the remains of an archegonium
are seen at _a_ (fig. 501, C) and the characteristic tent-pole apex
of the prothallus is shown at _b_. While in shape and in the general
plan of organisation _Rhabdospermum_ agrees with _Cardiocarpus_, the
vascular system in the chalazal region constitutes a distinguishing
feature. In _Rhabdospermum_ the main vascular strand passes through
the sclerotesta, _e_, _e′_, fig. 501, E, before giving off two bundles
which bend back (‘faisceaux récurrents’), traverse the shell, and then
pass up the sarcotesta in correspondence with the feebly developed
lateral keels as far as the micropyle, while in _Cardiocarpus_ (fig.
500, B) the bundles are given off before the main strand reaches the
sclerotesta. Similar recurrent bundles occur also in _Mitrospermum_
(fig. 494, K)[967].
=Mitrospermum.= A. Arber.
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