Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
Brongniart[1047] proposed this name for two species, _M. prismaticus_
and _M. difformis_, from French Coal Measures, but gave no description
of the specimens. The type-species, described from the Loire by
Grand’Eury as _Musocarpus prismaticus_[1048], is an ovate seed nearly
3 cm. long with six longitudinal ribs, three being more prominent
than the others, and characterised by a ribbed prolongation of
the base of uniform diameter spoken of as a carpophore, which was
apparently articulated to the lower part of the seed-proper from which
it was easily detached by a natural absciss-layer. Nothing is known
of the structure or of the affinity of the genus. Some specimens
in Dr Kidston’s collection from Westphalian beds in Lanarkshire
are closely allied to or perhaps identical with _M. prismaticus_.
In the presence of a distinct basal region _Musocarpus_ resembles
_Codonospermum_[1049], though without any knowledge of the anatomy of
the former type it is impossible to say whether or not this resemblance
has any morphological importance.
=Holcospermum.= Nathorst.
_Holcospermum sulcatum_ (Sternberg). The cast reproduced in fig. 506,
A, affords a good example of a form of seed recorded under several
generic names and not uncommon in Upper Palaeozoic rocks in Europe
and North America, which in most cases cannot be assigned to a genus
implying the possession of certain anatomical characters. This type
was figured by Sternberg[1050] from the Coal Measures of Radnitz as
_Carpolites sulcatus_. Some ‘fruits’ collected on the beach near
Newhaven, Midlothian, from the Calciferous Sandstone series, are
figured by Lindley and Hutton[1051] as _Carpolithes sulcata_, but
as already stated these have been transferred[1052] to the genus
_Rhynchogonium_. Several authors have referred specimens of the type
now included in _Holcospermum_ to _Rhabdocarpus_, but that genus, as
stated on a previous page, is restricted to seeds agreeing in form
with _R. tunicatus_. Renault[1053] instituted the genus _Colpospermum_
for a partially petrified seed from Commentry agreeing externally with
_Carpolites sulcatus_ Sternb. and regarded by him as specifically
identical, characterised by longitudinal ribs which represent folds
of the testa, the intervening grooves being occupied by an irregular
reticulum formed by occasionally anastomosing smaller ribs. The generic
name _Colpospermum_ should therefore be reserved for ribbed seeds
showing the anatomical features described by Renault and Zeiller:
its application to Sternberg’s species is inadvisable on the ground
that we have no information with regard to the morphological nature
of the ribbing. More recently Arber[1054] has proposed the name
_Platyspermum_, a name previously applied to a Cruciferous plant,
for Stephanian and Permian seeds formerly assigned to Berger’s genus
_Rhabdocarpus_, which are symmetrical in two planes. In this genus
he includes _Platyspermum sulcatum_ and among other species _P.
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