Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The difficulty of recognising the true nature of seed-like impressions
is illustrated by some specimens in the Goldenberg collection in
Stockholm described by Arber[188] as _Carpolithus Nathorsti_: these
consist of pieces of _Sphenopteris_ pinnae probably, as Zeiller
suggested, _Sphenopteris Schaumburg-Lippeana_ (Stur) bearing at
the ends of the segments of deeply divided pinnules what appeared
to be seeds 1 mm. long, oval and longitudinally ribbed, and
possibly enclosed in a cupule. Arber considered the ‘seeds’ to be
related to _Lagenostoma_, probably belonging to some member of the
Lyginopterideae. An examination of the specimens by Nathorst[189]
showed that the supposed seeds are collections of spores; but whether
the spores of a true Fern or the microspores of a Pteridosperm cannot
be determined.
_Pterispermostrobus_ Stopes.
_Pterispermostrobus bifurcatus_ Stopes.
Dr Stopes[190] has recently called attention to a resemblance between
specimens from the Westphalian of New Brunswick, described by her as
_Pterispermostrobus bifurcatus_, and _Lagenospermum Sinclairi_. The New
Brunswick fossil is made the type of a new genus _Pterispermostrobus_,
which is employed for fructifications of Pteridosperms that cannot
be associated with a known species of parent and may be either seeds
or complex male organs borne on a definitely branched rachis. The
type-species is represented by a slender axis bearing lateral branches
divided into two widely divergent arms each of which bears a terminal
body, 4 × 3 mm., characterised by 3–5 apical lobes extending 2 mm.
beyond the distal end of the seed-like organ and resembling a cupule.
In this as in many other cases it is impossible to determine the
true nature of the reproductive bodies, whether they are small seeds
or groups of microsporangia: the new generic name serves a useful
purpose though it is not always possible definitely to refer doubtful
fructifications of this kind to a Pteridosperm. The organs in question
may also be compared with _Codonotheca_[191].
_Pterispermostrobus pusillus_ (Nathorst).
The name, _Codonotheca pusilla_, is given by Nathorst[192] to some
doubtful specimens from the Culm of Spitzbergen representing short
stalks bearing linear-lanceolate scale-like bodies, 9–10 mm. long
by 1 mm. broad, coalescent at the base. Nathorst compares them
with Sellard’s species, _Codonotheca caduca_, but adds that they
may be cupules of some Pteridosperm and calls attention to their
resemblance to some fossils figured by Carpentier as _Calymmatotheca
acuta_. Both Nathorst’s species and the French specimens described
by Carpentier[193] as cupules may be referred to Dr Stopes’ genus
_Pterispermostrobus_ as their morphological nature cannot be determined.
+v.+ _Roots._
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