Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
In 1899 Zeiller[298] instituted the name _Potoniea_ for some peculiar
fertile leaves found in the Coal Measures of Heraclea consisting
of a branched axis with cuneate segments, 7–10 × 6–8 mm., bearing
numerous fusiform bodies, 1–1·5 mm. long, at the upper edge (fig.
421, A). These marginal bodies he regarded as sporangia and expressed
the opinion that _Potoniea_ may be the fructification of some form
of _Neuropteris_, _Mariopteris_ or _Alethopteris_. Carpentier[299]
afterwards described similar though rather larger examples from
the Pas-de-Calais coal-field as microsporophyll fragments of some
Pteridosperm: he stated that similar specimens had been found by
Kidston in England. In a later work Carpentier[300] described the
sporangia as crowded in groups (fig. 421, B) in the substance of the
thick lamina of _Potoniea_, and he connected the fertile segments
with _Neuropteris gigantea_, _N. pseudogigantea_ Pot. and _Linopteris
obliqua_ (Bunb.). Similar specimens are said to have been found in
Holland and Silesia. Bertrand[301] also records the association of
_Potoniea_ with _Neuropteris gigantea_ and _N. pseudogigantea_: he
describes some specimens as belonging to _N. gigantea_ Sternb. while
others, distinguished only by small differences, he attributes to _N.
pseudogigantea_. Kidston[302] has recently drawn attention to the
inconstancy of the characters mentioned by Potonié as distinguishing
features of _N. pseudogigantea_, and he shows good cause for referring
the examples so named to _N. gigantea_. The fertile lamina is almost
orbicular in surface-view and attached to a slightly excentric pedicel;
the microsporangia are borne on the lower surface and probably in
groups as described by Carpentier.
_Neuropteris Carpentieri_ Kidston. Kidston[303] has recently described
some fertile leaflets under this name from the Westphalian series of
South Staffordshire which he identifies with French examples referred
by Carpentier[304] to _Potoniea adiantiformis_ Zeill. The fertile
pinnules are thick and sub-cyclopteroid in form; the upper surface
bears densely packed, narrow and long, microsporangia, 4 × 0·5 mm.,
containing more or less spherical microspores 45–60μ in diameter;
the ventral face of the lamina on the removal of the spores shows
several strong veins. With these are associated sterile pinnules of
the Neuropteroid type, and examples are described intermediate between
the sterile and fertile leaflets. Kidston believes the specimens to be
microsporophylls of some species of _Neuropteris_, but as the material
does not suffice for identification with any known species a new name
is proposed. It is pointed out that in some states of preservation
the leaflets resemble Goeppert’s Permian species _Dictyothalamus
Schrollianus_[305].
_Alethopteris. Linopteris._
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