[Illustration: Fig. 429. Microspores of _Whittleseya elegans_. A, a group of spores; B, a single spore. (Preparations made from an American specimen, No. 2314, in Dr Kidston’s Collection.)] The specimens described by Lesquereux from Pennsylvania as _W. integrifolia_ and _W. undulata_ are less satisfactory than _W. elegans_. The Arkansas species _W. microphylla_[366], characterised by the obcuneate form of the lamina, is said to occur not only as detached leaflets but in loose bunches at the ends of slender axes, a circumstance favourable to the suggestion, based on the recently described English specimens, that the _Whittleseya_ leaves may be fertile pinnules of a Pteridosperm frond. Among other species attributed to Newberry’s genus is _W. brevifolia_ Wh. from Nova Scotia[367] with much smaller broadly triangular leaves 7 mm. long exclusive of the petiole and 8 mm. broad at the distal end (fig. 428, B). Dr Matthew[368] has also described a Canadian species _W. concinna_ from New Brunswick in beds assigned by Dr Stopes[369] to the Westphalian series. _Whittleseya fertilis_ Kidston. Since the discovery of _Whittleseya elegans_ in the Coal Measures of Staffordshire recorded by Mr Thomas, Dr Kidston has published an account of some specimens from the same district under the name _Whittleseya (?) fertilis_[370]: these consist of smaller cuneate scale-leaves or leaflets 1·4–2·4 cm. long and 8–9 mm. broad; the lamina has a dentate upper margin and is longitudinally ribbed. The scales occur in superposed pairs, closely fitting but not organically connected, at least in the state in which they are preserved; each pair forms a sporangium-like case enclosing numerous spores but the actual sporangia or synangia have not been preserved. Kidston describes the spores as 210–222μ in length, elliptical, and characterised in many cases by an oval slit; they are practically identical with the microspores of _W. elegans_. =DOLEROPHYLLUM=. Saporta.