indicates relatively drier conditions of climate than obtained when
perennial snow and ice covered wide areas of the low-ground goes
without saying, but I am unable to agree with those who maintain
that it implies a dry-as-dust climate, like that of some of the
steppe-regions of our own day. The remarkable commingling of arctic-
and steppe-faunas discovered in the Böhmer-Wald[AS] by Woldrich shows,
I think, that the jerboas, marmots, and hamster-rats were not incapable
of living in the same regions contemporaneously with lemmings,
arctic hares, Siberian social voles, etc. But when a cold epoch was
passing away the steppe-forms probably gradually replaced their
arctic congeners, as these migrated northwards during the continuous
amelioration of the climate.
[AS] Woldrich: _Sitzungsb. d. kais. Akad. d. W. math. nat. Cl._, 1880,
p. 7; 1881, p. 177; 1883, p. 978.
If the student of the Pleistocene faunas has certain advantages in the
fact that he has to deal with forms many of which are still living,
he labours at the same time under disadvantages which are unknown to
his colleagues who are engaged in the study of the life of far older
periods. The Pleistocene period was distinguished above all things
by its great oscillations of climate--the successive changes being
repeated and producing correlative migrations of floras and faunas.
We know that arctic and temperate faunas and floras flourished during
interglacial times, and a like succession of life-forms followed the
final disappearance of glacial conditions. A study of the organic
remains met with in any particular deposit will not necessarily,
therefore, enable us to assign these to their proper horizon. The
geographical position of the deposit, and its relation to Pleistocene
accumulations elsewhere, must clearly be taken into account. Already,
however, much has been done in this direction, and it is probable that
ere long we shall be able to arrive at a fair knowledge of the various
modifications which the Pleistocene floras and faunas experienced
during that protracted period of climatic changes of which I have been
speaking. We shall even possibly learn how often the arctic, steppe-,
prairie-, and forest-faunas, as they have been defined by Woldrich,
replaced each other. Even now some approximation to this better
knowledge has been made. Dr. Pohlig,[AT] for example, has compared the
remains of the Pleistocene faunas obtained at many different places
in Europe, and has presented us with a classification which, although
confessedly incomplete, yet serves to show the direction in which we
must look for further advances in this department of inquiry.
[AT] Pohlig: _Sitzungsb. d. niederrheinischen Gesellschaft zu Bonn_,
1884; _Zeitschr. d. deutsch. geolog. Ges._, 1887, p. 798. For a
very full account of the diluvial European and northern Asiatic
mammalian faunas by Woldrich, see _Mém. de l'Acad. des Sciences de St.
Pétersbourg_, vii^e sér., t. xxxv., 1887.
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