Adhesion between the tentacles and the prey is effected partly by the
gummy secretion of the glands of the ectoderm, which is perhaps
poisonous as well as adhesive, and partly by the threads. Once the prey
is fast and has ceased to struggle, it is brought to the mouth, which
opens wide to receive it, by the contraction and the contortions of the
tentacles, the column, and the peristome. At the same time a mass of
transparent mucus from the gastral cavity envelops it and assists in
dragging it in. There is some dispute as to the part played by the
tentacles in conveying food into the mouth. My own observations lead me
to think that, at any rate so far as _H. vulgaris_ is concerned, they do
not push it in, but sometimes in their contortions they even enter the
cavity accidentally.
When the food has once been engulfed some digestive fluid is apparently
poured out upon it. In _H. vulgaris_ it is retained in the upper part of
the cavity and the soluble parts are here dissolved out, the insoluble
parts such as the chitin of insect larvæ or crustacea being ejected from
the mouth. Digestion is, however, to a considerable extent
intracellular, for the cells of the endoderm have the power of thrusting
out from their surface lobular masses of their cell-substance in which
minute nutritive particles are enveloped and dissolved. The movements of
the cilia which can also be thrust out from and retracted into these
cells, keep the food in the gastral cavity in motion and probably turn
it round so as to expose all parts in turn to digestive action. Complete
digestion, at any rate in the Calcutta form, takes several days to
accomplish, and after the process is finished a flocculent mass of
colourless excreta is emitted from the mouth.
COLOUR.
In _Hydra viridis_, a species that has not yet been found in India, the
green colour is due to the presence in the cells of green corpuscles
which closely resemble those of the cells of certain freshwater sponges.
They represent a stage in the life-cycle of _Chlorella vulgaris_,
Beyerinck[AM], an alga which has been cultivated independently.
[Footnote AM: Bot. Zeitung, xlviii (1890): see p. 49, _antea_.]
In other species of the genus colour is largely dependent on food,
although minute corpuscles of a _dark_ green shade are sometimes found
in the cells of _H. oligactis_. In the Calcutta phase of _H. vulgaris_
colour is due entirely to amorphous particles situated mainly in the
cells of the endoderm. If the polyp is starved or exposed to a high
temperature, these particles disappear and it becomes practically
colourless. They probably form, therefore, some kind of food-reserve,
and it is noteworthy that a polyp kept in the unnatural conditions that
prevail in a small aquarium invariably becomes pale, and that its
excreta are not white and flocculent but contain dark granules
apparently identical with those found in the cells of coloured
individuals (p. 154).
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