Fungi: Their Nature and UsesCooke, M. C. (Mordecai Cubitt)
Science
Fungi: Their Nature and Uses
Cooke, M. C. (Mordecai Cubitt)
Fungi
[Illustration: FIG. 105.--Section of _Tubercularia_. _c._ Threads with
conidia.[K]]
Passing to the other bodies, which are of a deeper colour, it will
soon be discovered that, instead of being simple rounded heads, each
tubercle is composed of numerous smaller, nearly globose bodies,
closely packed together, often compressed, all united to a base
closely resembling the base of the other tubercles. If for a moment we
look at one of the tubercles near the spot where the crimson
tubercles seem to merge into the pink, we shall not only find them
particoloured, but that the red points are the identical globose
little heads just observed in clusters. This will lead to the
suspicion, which can afterwards be verified, that the red heads
are really produced on the stem or stroma of the pink tubercles.
A section of one of the red tubercles will show us how much the
internal structure differs. The little subglobose bodies which spring
from a common stroma or stem are hollow shells or capsules, externally
granular, internally filled with a gelatinous nucleus. They are,
indeed, the perithecia of a sphæriaceous fungus of the genus
_Nectria_, and the gelatinous nucleus contains the fructification.
Still further examination will show that this fructification consists
of cylindrical asci, each enclosing eight elliptical sporidia, closely
packed together, and mixed with slender threads called paraphyses.
Here, then, we have undoubted evidence of _Nectria cinnabarina_, with
its fruit, produced in asci growing from the stroma or stem, and in
intimate relationship with what was formerly named _Tubercularia
vulgaris_. A fungus with two forms of fruit, one proper to the pink,
or _Tubercularia_ form, with naked slender conidia, the other proper
to the mature fungus, enclosed in asci, and generated within the walls
of a perithecium. Instances of this kind are now known to be far from
uncommon, although they cannot always, or often, be so clearly and
distinctly traced as in the illustration which we have selected.
[Illustration: FIG. 106.--D. _Nectria_ surrounding _Tubercularia_; E.
tuft of _Nectria cinnabarina_; F. section of stroma; G. ascus and
paraphyses.]
It is not uncommon for the conidia of the _Sphæria_ to partake of the
characteristics of a mould, and then the perithecia are developed
amongst the conidial threads. A recently recorded instance of this
relates to _Sphæria Epochnii_, B. and Br.,[L] the conidia form of
which was long known before the _Sphæria_ related to it was
discovered, under the name of _Epochnium fungorum_. The _Epochnium_
forms a thin stratum, which overruns various species of _Corticium_.
The conidia are at first uniseptate. The perithecia of the _Sphæria_
are at first pale bottle-green, crowded in the centre of the
_Epochnium_, then black green granulated, sometimes depressed at the
summit, with a minute pore. The sporidia are strongly constricted in
the centre, at first uniseptate, with two nuclei in each division.
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