The facts are as follows. In all animals, when the appetite is very
intense, the stomach rises up, so that some people who have a clear
perception of this condition say that their stomach "creeps out" of
them; in others, who are still masticating their food and have not yet
worked it up properly in the mouth, the stomach obviously snatches
away the food from them against their will. In those animals,
therefore, which are naturally voracious, in whom the mouth cavity is
of generous proportions, and the stomach situated close to it (as in
the case of the synodont and channa), it is in no way surprising that,
when they are sufficiently hungry and are pursuing one of the smaller
animals, and are just on the point of catching it, the stomach should,
under the impulse of desire, spring into the mouth. And this cannot
possibly take place in any other way than by the stomach drawing the
food to itself by means of the gullet, as though by a hand. In fact,
just as we ourselves, in our eagerness to grasp more quickly something
lying before us, sometimes stretch out our whole bodies along with our
hands, so also the stomach stretches itself forward along with the
gullet, which is, as it were, its hand. And thus, in these animals in
whom those three factors co-exist--an excessive propensity for food, a
small gullet, and ample mouth proportions--in these, any slight
tendency to movement forwards brings the whole stomach into the mouth.
Now the constitution of the organs might itself suffice to give a
naturalist an indication of their functions. For Nature would never
have purposelessly constructed the oesophagus of two coats with
contrary dispositions; they must also have each been meant to have a
different action. The Erasistratean school, however, are capable of
anything rather than of recognizing the effects of Nature. Come,
therefore, let us demonstrate to them by animal dissection as well
that each of the two coats does exercise the activity which I have
stated. Take an animal, then; lay bare the structures surrounding the
gullet, without severing any of the nerves,[350] arteries, or veins
which are there situated; next divide with vertical incisions, from
the lower jaw to the thorax, the outer coat of the oesophagus (that
containing transverse fibres); then give the animal food and you will
see that it still swallows although the peristaltic function has been
abolished. If, again, in another animal, you cut through both
coats[351] with transverse incisions, you will observe that this
animal also swallows although the inner coat is no longer functioning.
From this it is clear that the animal can also swallow by either of
the two coats, although not so well as by both. For the following
also, in addition to other points, may be distinctly observed in the
dissection which I have described--that during deglutition the gullet
becomes slightly filled with air which is swallowed along with the
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