Grasses : $b A handbook for use in the field and laboratoryWard, H. Marshall (Harry Marshall)
Science
Grasses : $b A handbook for use in the field and laboratory
Ward, H. Marshall (Harry Marshall)
Grasses
[Illustration: Fig. 30. Diagram of a spikelet of a grass as it would
appear if the internodes between each set of organs were elongated.
_g^1_ lower and _g^2_ upper glume. _P_ lower and _p_ upper palea of the
second oldest flower _F^2_. _f_ a barren flower represented only by the
axis and paleæ. Above it a single palea and the termination of the axis
(_a_) of the spikelet.]
Inside them the axis or stalk (_rachis_) is continued for a short
distance only and on its sides are hinged two other pairs of more or
less boat-shaped scales, smaller and more delicate than the glumes, and
known as the _pales_ (_paleæ_), while a third pair of still smaller
pales is fixed to the end of the axis. In each case one smaller and
more delicate _inner_ palea is hinged just inside its more obvious
_outer_ palea. In the closed condition of the spikelet each of the
three pairs of pales is shut together, and pressed close to the axis,
and the pair of glumes shut in the whole.
On opening each of the lower pairs of pales we find a _flower_ inside;
but the terminal pair usually contain only the barren end of the axis.
Hence the latter is barren and the former are fertile.
Each fertile flower is found on careful dissection to consist of a
small swollen _Ovary_, or young grain, covered with silky hairs and
with a couple of delicate plumes (the _Stigma_) at its apex, and three
long and slender _Stamens_; while the magnifying glass will show two
tiny scales at the base--the _Lodicules_. All our ordinary grasses have
their flowers thus constructed--a pair of _lodicules_, three dangling
_stamens_ and an _ovary_ with a feathery two-plumed _stigma_: each such
flower is also enclosed in its pair of _pales_, and the several pairs
of pales of each spikelet, with their contents, are enclosed in the
pair of _glumes_ (Figs. 29-32).
Returning now to the inflorescence. It is clear that we have to
distinguish between the entire branched total Inflorescence, and the
Spikelets or partial inflorescences of which it is composed. In Botany
it is agreed to call any inflorescence consisting of a stalk or axis on
which the flowers are arranged without stalks--i.e. sitting directly
on it--and so that the youngest are above and the older below, a
_Spike_, and each spikelet is _a little spike_.
[Illustration: Fig. 31. Diagram of a spikelet of Wheat dissected (×
about 5) showing--from below upwards--the two glumes, two paleæ, two
lodicules, three stamens, and the ovary of the typical grass. Oliver.]
[Illustration: Fig. 32. Diagram of a spikelet of _Anthoxanthum_
dissected (× about 8), and showing--from below upwards--two outer and
two (awned) inner glumes, two paleæ, two stamens, and the ovary. There
are no lodicules. Oliver.]
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Elsewhere in the archive
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account