Grasses : $b A handbook for use in the field and laboratoryWard, H. Marshall (Harry Marshall)
Science
Grasses : $b A handbook for use in the field and laboratory
Ward, H. Marshall (Harry Marshall)
Grasses
On looking at the total inflorescence of the _Nardus_ we see that we
have a number of spikelets seated on the sides of an axis: this is then
a spike of spikelets, or, shortly, a _Spike_[9] (Fig. 5). _Mibora_ and
_Lepturus_ afford other examples. In _Panicum_, _Cynodon_ (Fig. 2) and
_Spartina_ we have groups of such spikes.
The _Poa_ inflorescence is, however, different. It consists of a loose
branched system of spikelets. Botanists term such a loosely branching
system, where each branch ends in a flower, a _panicle_: here then
we have a panicle of spikelets, or, shortly, a _Panicle_. _Aira_,
_Agrostis_, _Calamagrostis_, _Avena_, _Catabrosa_ (Fig. 4) and many
others afford further examples.
In _Dactylis_ we have a condition of affairs between the two extremes
given: the inflorescence is not so close a spike as _Nardus_, and
not so open a panicle as _Poa_--it is rather a spike-like panicle,
partaking of the nature of both. A special type of this (Foxtail)
occurring in certain grasses--e.g. _Phleum_, _Alopecurus_, _Phalaris_
and _Lagurus_,--is so characteristic as to be worth noting (Fig. 3).
There is also another aspect of these inflorescences which is not
without interest as showing how diagnostic characters may be obtained
from purely external features, easily observed in the field. We have
seen that in _Nardus_ the spikelets are arranged on one side only
of the rachis, or main axis, so that about three quarters of the
circumference of the latter is bare; whereas in _Lolium_--with which
_Agropyrum_ and _Brachypodium_ agree in this respect--the spikelets are
on opposite sides, leaving the intervening two quarters, i.e. half its
surface, of the circumference of the axis naked.
In _Cynosurus_ and the simpler forms of _Dactylis_, we find the
spikelets crowded round about three quarters of the surface of the
rachis, leaving the fourth quarter naked; and, finally, in _Phleum_,
_Alopecurus_, _Hordeum_, and _Anthoxanthum_ the spikelets cover the
entire surface.
In the first (_Nardus_) and third examples (_Cynosurus_, _Dactylis_)
where the spikelets are turned to one side, the inflorescence is said
to be _secund_.
Public-domain text, read in full here on John Shaqi.
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