Grasses : $b A handbook for use in the field and laboratoryWard, H. Marshall (Harry Marshall)
Science
Grasses : $b A handbook for use in the field and laboratory
Ward, H. Marshall (Harry Marshall)
Grasses
Returning to the floral diagram, we see that the two lodicules, the
three stamens and the ovary still remain to be explained. Much
discussion has been held regarding the lodicules. Functionally
they are said to aid in the divarication of the paleæ when the
period of anthesis arrives, and the stamens and stigmatic lobes
are to be exposed, by swelling and driving the valve-like paleæ
apart. Morphologically they have been explained as representing the
rudimentary perianth, here reduced to two minute scales, but in some
exotic grasses (_Bambusa_, _Stipa_, &c.) three lodicules, or even
more, are present. (Figs. 35, 36.) On the other hand they may be, and
probably are, scales of the nature of minute bracteoles and of no
significance to the flower itself.
[Illustration: Fig. 35. Floral diagram of ordinary grass. Each pair
of paleæ--_i.p._ inner and _o.p._ outer palea--encloses three stamens
(_s_), two lodicules (_l_) and the ovary. _st_, stigmatic plumes. _a_,
axis.]
[Illustration: Fig. 36. Floral diagram of a Bamboo, showing six
stamens, three inner (_i.s._) and three outer (_a.s._), and three
lodicules (_l_) in addition to the ovary. _i.p._ inner and _o.p._ outer
palea. _a_, axis.]
If this is so the flower of the grass is perfectly naked, and consists
in the typical case of three stamens and one carpel. The development
of the ovary lends no support to the view that there are two carpels:
the stigmatic plumes are not separate styles. Nor does the fact that
some grasses have six stamens lend support to the idea that the flower
is derived from the trimerous type so common in Monocotyledons: other
numbers may occur--e.g. as few as two (_Anthoxanthum_, Fig. 32) or even
one only (_Uniola_, _Cinna_, &c.) or as many as 20 or 40 in certain
other exotic grasses.
Even when three stigmatic plumes are developed, as in some Bamboos,
close investigation does not confirm the view that the ovary consists
of more than one carpel.
We must therefore regard the flower of the grass as typically composed
of one carpel and three stamens, with no perianth whatever. It is
subtended by one or more bracteoles (the lodicules), and enclosed in a
pair of bracteoles one higher than the other (the paleæ). The glumes
are bracts of the partial inflorescence--the spikelet.
That there are some departures from this type in detail does not
invalidate the importance of the fact that most grasses conform to it.
I now pass to the consideration of a grouping of our ordinary grasses
according to their floral arrangements.
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