Grasses : $b A handbook for use in the field and laboratoryWard, H. Marshall (Harry Marshall)
Science
Grasses : $b A handbook for use in the field and laboratory
Ward, H. Marshall (Harry Marshall)
Grasses
_x Palea distinctly bifid at the apex: awn nearly
twice its length. A very small grass about six
inches high: leaves subulate, fine and short._
_A. caryophyllea_, L.
_xx Paleæ almost imperceptibly bidentate: awn not much
longer than palea. About 12-18 inches high: leaves
inrolled._
_A. flexuosa_, L.
Other species of _Aira_, with more condensed inflorescences, are
dealt with on p. 112. _Agrostis_ is distinguished by the leaves and
one-flowered spikelets (see p. 104). _Poa_ and _Catabrosa_ have no
awns.
CHAPTER VIII.
THE FRUIT AND SEED.
The stigma of an ordinary grass consists of two divaricating plume-like
structures composed of thin-walled cells. When the paleæ open these
stigmatic plumes protrude, one on either side, and readily catch pollen
shed from the dangling stamens and carried by the wind, and since the
pollen of the same flower is usually shed at a time when the stigmas
of many neighbouring plants are mature, there is every opportunity for
cross-fertilisation. (Fig. 33.)
In some cases, however, e.g. _Anthoxanthum_, _Alopecurus_, the flowers
are proterogynous, the stigmatic plumes being ready for pollination
some time before the pollen is shed from the anthers of the same
flower; whereas in most of our grasses the pollen begins to scatter
before the stigmas are ready (protandrous). Among exotic grasses, many
are diœcious or monœcious--i.e. the flowers contain stamens
only or ovary only, on each plant, or on different inflorescences
of the same plant respectively--and even in our own _Holcus_ and
_Arrhenatherum_ this state of affairs is partially represented, since
one flower of the spikelet is male only.
In some grasses, e.g. Rye, however, it appears improbable that
cross-fertilisation ever occurs, since the paleæ do not open, and the
pollen falls on to the stigma direct; and in _Leersia_ and the foreign
_Amphicarpum_ the spikelets are completely cleistogamous, those of the
latter being on stalks close to the ground which push the flowers into
the soil, where pollination and fructification are accomplished.
Hybrid grasses are by no means uncommon. To say nothing of the numerous
cross-bred Wheats and Barleys, artificial hybrids have been raised
between Wheat and Rye. In the Maize an astonishing number of selected
cross-breeds have been obtained, and, among others, certain forms in
which the seeds have a violet outer layer and a sugary endosperm, are
found to transmit these characters to the resulting seed of a variety
which would normally have produced seed with white outer layers and
starchy endosperm, if the pollen of the former is used on the stigma of
the latter. Such direct influences of the pollen are termed _Xenia_.
Public-domain text, read in full here on John Shaqi.
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