Histology of medicinal plantsMansfield, William James
Science
Histology of medicinal plants
Mansfield, William James
Botany, Medical; Plant anatomy
STEMS
When studying stems it should first be determined whether they were
derived from monocotyledonous or dicotyledonous plants. This fact is
ascertained by determining the type of the fibro-vascular bundle. See
Chapter XI. The next fact to determine is whether the stem is from an
herb or from a woody plant. This fact is readily determined because
herbaceous stems have a true epidermis, masses of collenchyma at
the angles of the stem. The cortical cells contain chlorophyll, and
the pith is very large. Woody stems have a corky layer, a phellogen
layer, and the pith is very small except in the very young woody
stems.
Having determined these facts, a study should be made of the
arrangement, form, structure, color, and the cell contents of the
different cells in order to determine the species of plant from which
the stem was obtained.
HERBACEOUS STEMS
The great variation in the structure of herbaceous stems is shown
in the cross-sections of spigelia (Plate 95); in ruellia (Plate
96); in the charts of powdered genuine horehound, powdered spurious
horehound, and in the chart of powdered insect flower stems.
CROSS-SECTION SPIGELIA STEM
Spigelia stem (Plate 95) has the following characteristic structure:
=Epidermis.= The epidermal cells are papillate.
=Cortex.= The cortical parenchyma cells consist of tangentially
elongated cells which are oval in outline.
=Phloem.= The phloem consists of sieve cells, phloem parenchyma, and
of bast fibres.
[Illustration: PLATE 95
CROSS-SECTION OF STEM OF SPIGELIA MARYLANDICA, L.
1. Papillate epidermis. 4. Phloem. 7. Inner phloem.
2. Cortical parenchyma. 5. Cambium. 8. Pith.
3. Bast stereome. 6. Xylem.]
The =sieve cells= are small, and with thin, white, angled walls.
The =phloem parenchyma= resembles the sieve cells, but they are
larger.
The bast fibres are rounded in outline and the walls are thick,
white, non-porous, and non-striated.
=Cambium.= The cambium cells are rectangular in shape or the walls
are collapsed and the cells indistinct.
=Xylem.= The xylem contains vessels, wood parenchyma, medullary rays.
The vessels are small and angled, the walls are thick and white.
=Wood parenchyma.= The cells are variable in size and shape, and
the walls are thick. The medullary ray cells are small, narrow, and
tangentially elongated.
=Internal Phloem.= External to the pith parenchyma are isolated
groups of internal phloem consisting of sieve cells.
=Pith Parenchyma.= The pith parenchyma cells are oval in form and
irregularly placed. The cells contain small, simple starch grains.
RUELLIA STEM
The cross-section of ruellia stem (Plate 96) is as follows:
=Epidermis.= The epidermal cells are variable in shape and very
large. There are no cell contents.
=Cortex.= The cortex consists of collenchyma and parenchyma cells and
stone cells.
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