Histology of medicinal plantsMansfield, William James
Science
Histology of medicinal plants
Mansfield, William James
Botany, Medical; Plant anatomy
There are three well-defined types of the fibro-vascular bundle,
namely, the =radial=, the =concentric=, and the =collateral= types.
RADIAL VASCULAR BUNDLES
The radial type of bundle is met with most frequently in
monocotyledonous roots.
In this form (Plate 114) the xylem forms radial bands of tissue
which alternate with isolated groups of phloem. The space between
the phloem and xylem is filled in with either parenchyma or fibres,
or both. In some cases the vessels of the xylem meet in the centre
of the root, while in other cases the centre of the stem is occupied
by pith parenchyma. Each bundle is surrounded by parenchyma cells,
and in iris, calamus, and veratrum, rhizomes, and endodermis,
surrounds the bundles located in the centre of the stem, consisting
of thin-walled (mechanical) cells.
[Illustration: PLATE 124
CROSS-SECTION OF A RADIAL VASCULAR BUNDLE OF SKUNK CABBAGE ROOT
(_Symplocarpus fœtidus_ [L.], Nutt.)
1. Vessels.
2. Bundle sheath.
3. Parenchyma.
4. Sieve cells.]
[Illustration: PLATE 125
CROSS-SECTION OF A PHLOEM-CENTRIC BUNDLE OF CALAMUS RHIZOME
(_Acorus calamus_, L.)
1. Vessels.
2. Sieve cells.
3. Phloem parenchyma.
4. Parenchyma surrounding the bundles.]
In sarsaparilla root, the pith is composed of thick-walled, porous
pith parenchyma cells with starch. Outside the pith are arranged
radial bands of oval vessels which decrease in size toward the
periphery. Between the ends of these bands occur isolated groups of
sieve cells.
Surrounding the sieve cells and vessels are thick-walled, angled
fibres.
External to these cells is an endodermis composed of lignified
brownish-colored cells one layer in thickness.
CONCENTRIC VASCULAR BUNDLES
There are two principal types of the concentric bundle, namely,
xylem-centric, in which the xylem is centric and the phloem is
peripheral, as in veratrum root; and phloem-centric (Plate 125), in
which the phloem is centric and the xylem peripheral, as in calamus
rhizome.
COLLATERAL VASCULAR BUNDLES
There are three types of collateral vascular bundles--namely, closed
collateral, bi-collateral, and open collateral.
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