Histology of medicinal plantsMansfield, William James
Science
Histology of medicinal plants
Mansfield, William James
Botany, Medical; Plant anatomy
The =relation of the stoma to surrounding cells= is best shown in
cross-sections of the leaf. In powders the relationship of the stoma
to the surrounding cells is, however, readily ascertained. If the
guard cells come in focus first, they are above the surface; if the
guard cells and the surrounding cells come in focus at the same time,
the stomata are even with the surface; if the stomata come in focus
after the surrounding cells, they are below the surface of the leaf.
The relationship of the stoma to the surrounding cells should always
be ascertained, not only in cross-sections of the leaf, but also in
powders.
There is the greatest possible variation in the size of guard cells.
This fact must always be kept in mind when studying leaves. This
variation in the size of the guard cells is clearly illustrated by
coca, senna, and by deer’s-tongue. In coca the stomata are very
small; in senna they are larger; while in deer’s-tongue the stomata
are very large.
The width and length of the stoma or opening between the guard cells
are of a character which must not be overlooked. Generally speaking,
those leaves which have large guard cells will have correspondingly
large stomata.
The guard cells usually contain chloroplasts showing various stages
of decomposition.
In bay-rum leaf the guard cells are of a bright reddish-brown color,
but in most leaves the guard cells are colorless.
LENTICELS
=Lenticels= are small openings occurring in the bark of plants.
The lenticels bear the same relationship to the stem that the
stomata do to the leaves. Lenticels, like stomata, have a threefold
function--namely, exchange of gases in photosynthesis, in
respiration, and the giving off of water.
Lenticels are macroscopically as well as microscopically important.
When unmagnified the lenticels are circular, lens-shaped, or
irregular in outline. They are arranged in parallel longitudinal
lines or parallel transverse lines, or they are irregularly
scattered. The latter is the usual arrangement. In most cases they
are elevated slightly above the surface of the bark. In root barks
particularly the lenticels stand out prominently from the surface of
the bark and in many cases appear stalked.
The color of the lenticels differs greatly in the different plants.
In acer spicatium they are brown; in witch-hazel they are gray; in
xanthoxylium they are yellowish; and lastly, the number of lenticels
occurring in a given surface of the bark should always be considered.
On cross-sections the lenticel (Plate 57, Fig. 2) is seen to have
a central depressed portion made up of loosely arranged cells.
Bordering the cavity are typical cork cells. The cork cells
immediately surrounding the lenticels are usually darker in color,
and many of the cells are partly broken down.
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