Histology of medicinal plantsMansfield, William James
Science
Histology of medicinal plants
Mansfield, William James
Botany, Medical; Plant anatomy
The =chloroplasts= of the higher plants are green, and they vary
somewhat in size, but they have a similar structure and form.
Chloroplasts are mostly oval in longitudinal view and rounded in
cross-section view. Each chlorophyll grain has an extremely thin
outer wall, which encloses the protoplasmic substance, the green
granules, a green pigment (chlorophyll), and a yellow pigment
(xanthophyll). Frequently the wall includes starch, oil drops, and
protein crystals.
Chloroplasts are arranged either in a regular peripheral manner along
the walls, or they are diffused throughout the protoplast.
The palisade cells of most leaves are packed with chlorophyll grains.
In the mesophyll cells the chlorophyll grains are not so numerous,
and they are arranged peripherally around the innermost part of the
wall.
Chloroplasts multiply by fission--that is, each chloroplast divides
into two equal halves, each of which develops into a normal
chloroplast.
Chlorophyll occurs in the palisade, spongy parenchyma, and guard
cells of the leaf; in the collenchyma and parenchyma of the cortex
of the stems of herbs and of young woody stems, and, under certain
conditions, in rhizomes and roots exposed to light. Almost without
exception young seeds and fruits have chlorophyll.
In powdered leaves, stems, etc., the chlorophyll grains occur in the
cells as greenish, more or less structureless masses. Yet cells with
chlorophyll are readily distinguished from cells with other cell
contents. In witch-hazel leaf the chlorophyll grains appear brownish
in color. Powdered leaves and herbs are readily distinguished from
bark, wood, root, and flower powders.
Leaves and the stems of herbs are of a bright-green color. With the
exception of the guard cells, the chloroplasts occur one or more
layers below the epidermis; but, owing to the translucent nature of
the outer walls of these cells, the outer cells of leaves and stems
appear green.
Wild cherry, sweet birch, and, in fact, most trees with smooth barks
have =chloroplasts= in several of the outer layers of the cortical
parenchyma. When the thin outer bark is removed from these plants,
the underlying layers are seen to be of a bright-green color.
LEUCOPLASTIDS
=Leucoplastids=, or colorless plastids, occur in the underground
portions of the plant; they may, when these organs in which they
occur are exposed to light, change to chloroplastids.
Leucoplasts are the builders of starch grains. They take the chemical
substance starch and build or mould it into starch grains, storage
starch, or reserve starch.
Other characteristic chromoplasts found in plants are yellow and
red. Yellow chromoplasts occur in carrot root and nasturtium flower
petals. Red plastids occur in the ripe fruit of capsicum.
STARCH GRAINS
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