Histology of medicinal plantsMansfield, William James
Science
Histology of medicinal plants
Mansfield, William James
Botany, Medical; Plant anatomy
Other starches with rounded surfaces are shown on Plates 72 and 73.
Angled outlined grains are common to cardamon seed, white pepper,
cubebs, grains of paradise (Plate 76, Fig. 4), and to corn (Plate 70,
Fig. 3).
The outlines of all compound grains are made up partly of plane and
partly of curved surfaces.
SIZE
The =size= (greatest diameter) of starch varies greatly even in the
same species, but for each plant there is a normal variation.
In spherical starch grains the size of the individual grains is
invariable, but in elongated starch grains and in parts of compound
grains the size will vary according to the part of the grain
measured. In zedoary starch (Plate 71, Fig. 4), for instance, the
size will vary according to whether the end, side, or surface of the
starch grain is in focus.
The parts of compound grains often vary greatly in size. Such a
variation is shown in Plate 75, Fig. 2.
HILUM
The =hilum= is the starting-point of the starch grain or the first
part of the grain laid down by the amyloplast. The hilum will be
central if formed in the middle of the amyloplast, and excentral if
formed near the surface of the amyloplast. It has been shown that the
developing starch grain with eccentric hilum usually extends the wall
of the amyloplast if it does not actually break through the wall.
Starch grains with excentral hilums are therefore longer than broad.
[Illustration: PLATE 70
STARCH
1. Calabar bean (_Physostigma venenosum_, Balfour).
2. Marshmallow root (_Althæa officinalis_, L.).
3. Field corn (_Zea mays_, L.).]
[Illustration: PLATE 71
STARCH
1. Galanga root (_Alpinia officinarum_, Hance). 2. Kola nut (_Cola
vera_, [K.] Schum.). 3. Geranium rhizome (_Geranium maculatum_ L.).
4. Zedoary root (_Curcuma zedoaria_, Rosc.). 4-_A_. Surface view of
starch grain. 4-_B_. Side view of starch grain. 4-_C_. End view of
starch grain.]
In central hilum starch grains the grain is laid down around the
hilum in the form of concentric layers. These layers are of variable
density. The dense layers are formed when plenty of sugar is
available, and the less dense layers are formed when little sugar
is available. The unequal density of the different layers gives the
striated appearance characteristic of so many starch grains.
In eccentric hilum starch grains the starch will be deposited in
layers which are outside of and successively farther from the hilum.
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